Tracking the evolution of Joey and Shooby’s impossibly pure bromance on The Circle – The A.V. Club

On its surface, The Circlesounds like some shallow, bingeable trash. Stick around until the second episode, however, and youll likely be hooked. The social media competition that forms the spine of the Netflix seriesplayers communicate solely through a social media platform, trying to convey authenticity so as not to get blocked by the games influencersisnt all that gripping in a vacuum, but its reliance on the personalities and machinations of its cast is the key to its appeal. Shockingly, even the most duplicitous of the contestantssome purposely catfish their colleaguesturn out to be compelling, amiable folks, each working to convey genuine emotions through a deceptive avatar. But, as the show makes abundantly clear, ones genuine self is capable ofbleeding through a chat box. Four of the five finalists, after all, survived the entirety of the game without once masking their identity or motivations. The friendships they made really felt real.

Just look at Joey Sasso and Shubham Goel. The former is a buff, flirtatious bartender with a thick Jersey accent while the latter is a socially awkward, Marvel-loving virtual reality engineertogether, however, they formed one of the sweetest, most soul-affirming bonds on TV. Their friendship started simply, with Joey observing in the second episode that Shubhama.k.a. Shoobywas buggin last night and Shooby fretting over whether adding man to the end of a casual inquiry sounded weird. Now, with the show behind them, the pair have made it clear their bromance will persevere beyond the cameras.

In every way possible, we are different, but thats the beauty of it, Shooby recently told Cosmo. He was my day-one guy in there. Me and Joey still keep in touch....When Im back [in Los Angeles], were definitely gonna kick it. Im hoping to kick it, like, three or four times a week. With Joey, I one hundred percent know that our relationship from that Circle is going to be just as strong outside of it.

So, to ensure we never forget the exquisite nature of their union, please join us for a pictorial trek through their friendships most pivotal moments.

(Note: I will be spelling it Shooby, despite Netflixs closed captioning spelling it Shubby. It sounds like Shooby, guys.)

#FriendsTillTheEnd, indeed.

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"Expanding the boundaries of lighting" The evolution of lighting as a platform to create enhanced retail experiences – Retail Dive

Light has always had the ability to transform spaces, so as the leading lighting manufacturer, we are compelled to help shape the future of illumination. Considering a future state where digital lighting is intelligent and connected has led Acuity Brands to expand from simply illuminating a space to empowering lighting to solve retail customers' challenges in their stores.

Our corporate commitment and vision of Expanding the boundaries of lightingmeans developing advanced lighting and controls products that are foundational to supporting new and connected IoT solutions. One of the ways we do this is by embedding lighting with digital technologies such as controls, networking devices and sensors to form a platform for location-aware service. We call this a sensory network at Acuity Brands, which is powered and present where people shop and work. We also apply our deep experience in lighting manufacturing to bring this technology to scale, opening applicability beyond high-value retail sales floors into warehouses, manufacturing, and healthcare spaces.

How do we imagine this future of lighting? By listening to our customers. Acuity Brands responds to the needs of retail customers who seek to improve the in-store shopping experience. In addition, we are actively engaged in transforming store operations through real time technologies that improve associate efficiency and support ship-from-store and buy online/pickup at store services. Digital transformation is happening in brick and mortar retail with the help of accurate indoor location-awareness powered by Acuity Brands.

Now that lighting goes beyond energy savings provided by LEDs to form an IoT platform, we connect with building management, refrigeration, and other devices to harvest data that informs facility teams where and how to improve operations with the biggest impact. Our innovations provide analytics and insights on sensor data delivered through the lighting network.

At the start of a major renovation, retailers look for the best solutions in design and materials specifications for LED luminaires and lighting controls. Learning more about the Atrius sensory network from Acuity Brands, the conversation quickly moves to the additional capabilities of the platform. The LED luminaires embedded with BLE (Bluetooth Low Energy) assist visitors with wayfinding and deliver location awareness to tags and monitor assets throughout the building space.

When coupled with low-power sensors, the lighting network can monitor customer foot traffic or alert store associates to items that need to be restocked. These are simple use cases expanding the purpose of lighting: providing a deeper understanding of customers and their in-store shopping experience through rich data insights. At the risk of sounding clich, these conversations create "lightbulb moments" for the retailer.

According to research firm Forrester, an estimated 27% of global retailers and wholesalers indicate that increasing usage of data insights is a high priority in 2020. However, 38% of those surveyed said, "improving their use of data insights in business decision making will be very or even extremely challenging". This has led Forrester to predict that while "data mastery will be the common thread among retail winners in 2020 few will achieve it" .

Addressing this challenge, Acuity Brands brings experience and leadership to tackle critical insights. Walgreens, Target and other successful retailers with Acuity Brands digitally connected lighting systems installed are now activating additional features and functionality to capture data insights that give their business the information needed to improve customer experience by understanding customer preferences and identifying opportunities for improvement.

In the case of Walgreens, their Acuity lighting network was the first layer of implementation. Through a partnership with Microsoft, connected capabilities such as Atrius Retail Solutions were implemented to capture data to enhance the customer journey and get Walgreens closer to their vision to improve the overall shopper experience.

By creating solutions for retailers that deliver business insights from the connected lighting network, renovation investments become-technology investments and have the capacity to serve the business well into the future as highlighted in the recent Walgreens News that covers their first year of partnering with Microsoft on the "future of healthcare" initiative.

Leveraging data, many of our customers are learning new insights such as shopper in-store foot traffic patterns and improving loss prevention. By implementing Atrius Retail Solutions, they gain actionable information that significantly impacts their business. Distilling the panoply of harvested data to action is another way we're expanding the purpose of lighting.

Visit the Atrius website to learn how today's networked lighting solutions can help solve business challenges with practical data-driven solutions through Atrius Retail Solutions.

Download the Navigant white paperClosing the Online/Offline Gap for Retail Storessponsored by Acuity Brands.

Fiona Swerdlow,Predictions 2020: Retailers Seek An Edge Amid Uncertainty, Forrester Research

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Maple Leafs Mitch Marner on first All-Star Game, evolution of NHL wingers – Sportsnet.ca

After a pair of impressive 60-point campaigns as a young gun and a dominant 94-point effort last season, Mitch Marner is officially an All-Star.

The star winger earned his first career selection on the back of the Maple Leafs faithful, his name added to the mix via the leagues Last Men In fan vote.

With an impressive 47 points on the season so far, second-most on the team despite missing roughly a month with an early injury, the 22-year-old said hes honoured to get his first shot at joining the All-Star festivities.

Its definitely up there, Marner told Sportsnet 590s Hockey Central crew when asked where an All-Star debut ranks among young hockey players goals. I mean, probably top three Id say, [with] the Stanley Cup, getting drafted and then making it here. Its something as a kid you watch, you grew up watching. You watched all the guys competing and just really looking like theyre having a blast.

Im super honoured to be a part of it now.

The Markham, Ont., native will suit up for the inaugural Shooting Stars event as part of the Skills Competition on Friday, a trick-shot-like affair thatll see players wiring pucks off an elevated platform 30 feet above the ice, trying to snipe targets spread across the sheet.

We went and gave it a shot yesterday from where the actual mark was, where everythings sitting in the stands, Marner said. Its weird I mean, youre elevated, you dont really know how much power to put into it or anything, but well see how it goes. Hopefully, I just dont embarrass myself.

Hockey Central

Mitch Marner will also be in the stands this All-Star weekend

January 24 2020

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The skill-set Marners put on display so far in his young career suggests hell be just fine. But theres far more to his game and that of his fellow All-Star wingers than the offence-focused skill-sets of the stars who played his position back in the day.

The evolution of the game has seemed to bring with it a blurring of the lines for players in all positions. And while the focus of that transition is most often the greater offensive roles of todays top blue-liners, theres been just as big a shift for the wingers, says Marner.

I think for wingers, you see more now the defensive wingers or two-way wingers that have really developed throughout the league, Mark Stone being the kind of lead guy in that, Marner told the Hockey Central crew. Seeing what hes done the last couple years with takeaways and even just with his stick, hes been amazing at that. So, I think really the winger role has evolved into being something bigger than just kind of that guy that goes up and down the wing, that can shoot, that power forward guy getting the puck deep and hitting guys.

I think its really developed a lot now into a skilled guy, a playmaker or a goal-scorer or really anything. I think now in wingers minds its not just about doing that up-and-down ice, its about covering all zones and making sure youre covering for your centreman if need be, and I think everyones kind of taken a bigger role with that.

Listen to Mitch Marners full interview with Sportsnet 590s Hockey Central via the audio player embedded within this post.

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A top UiPath exec called its recent layoffs a ‘natural part of the evolution.’ Here’s how the hot AI startup now plans to scale in 2020. – Business…

BELLEVUE, WASHINGTON At the start of 2019, UiPath announced a $568 million funding round valuing the startup at $7 billion.

It quickly became the world's most-funded AI startups and was hailed by analysts as nearly unbeatable in its offerings. The company specializes in robotic process automation, which aims to help companies automate common, repetitive, and mundane computer tasks.

Then, in October, UiPath laid off roughly 400 jobs and parted ways with a chief financial officer who had only been at the company since January. The move was aimed at eliminating positions or individuals that were key to achieving success as a startup, but were not suited to help UiPath continue to grow, according to head of AI products PD Singh.

Instead, the firm had to prepare for its future including a potential initial public offering (or IPO) and that required individuals with a growth mindset. PD Singh is the head of AI products at UiPath UiPath

"It was just a natural part of that evolution that you needed to get rid of that baggage," he told Business Insider at UiPath's Bellevue, Washington, office. "These things are natural. This was a startup which was working in a very scrappy mode but that's not how things eventually would be done in a company which is either IPO'ing or post-IPO."

Singh did not expect any additional layoffs in 2020, but noted that anything was possible. "Everybody is dispensable here, including me," he said. "If there's one thing I've learned in technology, you shouldn't get hung up about layoffs."

Instead, UiPath is looking to put those layoffs behind them and scale the business in 2020. Last year, "we put good stakes in the ground in terms of the product lanes that are needed by our customers," Singh said, adding that this year will be all about "scaling the installation of those product lines."

Singh a former head of product at Microsoft who joined UiPath in 2018 outlined the three steps the company is taking to grow.

RPA has the potential to save companies hundreds of millions of dollars every year in reduced operating costs. But for UiPath to grow, Singh says it needs to find more real-world examples to tout.

"With RPA being a fledgling industry and AI itself being a fledgling industry one of the things that's going to make us not be too buzzy or not be too frothy is real use cases. And real use cases working at scale," he said.

UiPath already has big-name clients using its platform. Chevron, for example, is using RPA in a limited capacity to automatically open email attachments and upload them to a common database, a project they want to scale to the whole enterprise.

Japan's Sumitomo Mitsui Banking Corporation is also using the technology to save an estimated $500 million annually by the end of 2020.

Similar to how developers helped populate the app store for iPhone, UiPath is hoping to expand its network of partners that can build upon its own platform.

"Having that partner network is super important," said Singh. "In the AI team right now, I have [an] initiative to light up our strategic [global systems integrators] that we will train" to use UiPath's offerings to develop custom solutions for clients.

That goal is relatively easy, he added, because the company can focus on high-demand verticals based upon how quickly the industries can adopt the tech.

Healthcare, for example, is much more highly regulated and has a higher barrier of entry for new applications. On the other hand, oil and gas companies can typically implement technology much faster.

Singh is confident in UiPath's current product line, but says a goal will be to continue meeting customer demands for new offerings.

"If you have a good sense of what your customers want and you keep innovating in the direction of the customer need, that's what makes your company the top company in the world," he said.

UiPath gets ample feedback from clients to help inform future development. A key metric is daily active robots, which simply refers to how many models are currently in use. The company is also active on forums with developers to learn about pain points with its tools, and it offers select customers private previews of offerings not yet ready for primetime.

"We have about 150 different models that are in production," Singh said.

If you are an individual who was laid off from UiPath (or a current employee) and would like to share your experience at the company, I can be reached at JWilliams@businessinsider.com or on Signal/WhatsApp at (309) 265-6120.

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A top UiPath exec called its recent layoffs a 'natural part of the evolution.' Here's how the hot AI startup now plans to scale in 2020. - Business...

5G Will Bring Forth A Cryptocurrency Evolution In Payments Within The Next Half-Decade – The Coin Republic

Steve Anderrson Friday, 24 January 2020, 06:23 EST Modified date: Friday, 24 January 2020, 06:23 EST

The blazing speeds and more efficient communications protocols that 5G expected to bring in expected to change the face of cryptocurrency-based payments in the average consumer space within the next five years.

It presumed that within 2025, nearly half of all smartphone owners who do not have access to a bank or a financial institution would be able to make cryptocurrency transactions for their daily business needs.

Managing VP at Gartner, Miriam Burt shared the same opinions at the Gartners Strategic Predictions for 2020 and Beyond keynote. It held at the Oracle OpenWorld 2020, which was hosted at the Dubai Trade Centre Arena this week.

She went on to state her belief that for financial leaders to remain competitive and have full customer satisfaction, the future of finance based on cryptocurrencies must undoubtedly be thought of and considered.

Were looking at the age of fintech, where there is a potential for all payments to be cashless and make financial services accessible to a whole new population that does not have access to banking services.

Burt went on to say that

In the Middle East and North Africa (MENA) region, two-thirds of people have no bank accounts; however, more than half of the countries have no bank accounts. However, more than half of the countries have a mobile subscriber penetration rate of 70% of the population. This means that we have more people who have phones than bank accounts. An estimated 23% of these people are under-banked, so they have access to smartphones but dont have access to traditional financial services.

What Burt is trying to convey is the idea that the onset of cryptocurrencies will bring financial services one step closer to people without having to go through complicated procedures that take time and infrastructure to build and maintain.

The phones are already in place, in the hands of the people, and the bases for the financial services are already there too within cryptocurrencies.

Cryptocurrencies will become the core of mobile payments, and digital payments in general, and bring financial institutions closer to the people globally.

Burt also said that China Telecom would be the first to implement this technology, and they are already working on blockchain technology-based projects that will use 5G SIM cards to transact cryptocurrencies.

Successful implementation of the above initiative could put China Telecom as a massive lead in telecommunications over other countries, and it will become highly popular.

They will have extremely secure standards, including next-level digital authentication and decentralized authentication, to allow cryptocurrency payments.

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5G Will Bring Forth A Cryptocurrency Evolution In Payments Within The Next Half-Decade - The Coin Republic

Scientific Decadence and the Myth of Objectivity – Discovery Institute

Writing here yesterday about the mischaracterization of 437-million-year-old scorpion fossils, paleontologist Gnter Bechly used an apt but unexpected word to describe evolutionary thinking. It isnt just mistaken. It reflects a state of decadence:

In todays science world it is no longer sufficient to objectively describe some nicely preserved ancient fossils. You must overinterpret the evidence and oversell their importance with a fancy evolutionary narrative. And you do not have to hesitate to be really bold with your claims, because neither the scientific reviewers nor the popular science media will care if your claims are actually supported by the evidence. This system is broken. It was broken by the pressure to publish or perish, by the pressure of public relation departments to generate lurid headlines, and by the pressure of the idiotic paradigm that nothing makes sense in biology except in the light of evolution.

Under these circumstances, Good science falls by the wayside. Now neuroscientist Michael Egnor comments over at Mind Matters on decadence in a completely different scientific field: medicine and medical ethics. Abortion advocates have long advanced the falsehood that babies in the womb feel less pain than we do or no pain at all. The truth is the exact opposite: An unborn child with an immature brain probably experiences painmoreintensely than an individual with a mature cortex.

Perhaps the most disturbing damage that the abortion lobby has done to our society aside from the systematic killing of tens of millions of innocent human beings is the corruption of science in the name of ideology. Nowhere is this corruption more obvious than in the misrepresentation of the neuroscience of fetal pain perception.

A new article in theJournal of Medical EthicstitledReconsidering Fetal Pain(open access)is a welcome correction to the abortion lobbys systematic misrepresentation. The authors, one of whom is an abortionadvocate,reviewed the literature on the perception of fetal pain and came to the conclusion that there is clear scientific evidence to support the view that unborn children feel pain as early as 13 weeks of gestation.

This should not have come as a surprise, since doctors who work with newborns and premature babies routinely observe that they respond with screams to a needle prick that an adult would barely register.

Dr. Bechly calls it decadence. Dr. Egnor calls it corruption. Its one and the same thing: whereas, according to widespread legend, scientists just objectively sift facts, in reality ideological ax-grinding is common and probably worse than it ever was. Remember, what Bechly and Egnor are describing isnt limited to a stray scientist here or there. It is systematic. Hence the state of decadence.

As the new Long Story Short video from Discovery Institute on homology puts it, Scientists are just like everyone else: people. And we can be uncritical of things that we want to believe.

Thats a charitable way of putting it. In one way, scientists arent like everyone else: that is, because of the enormous prestige they enjoy, the impact of their being uncritical of things that [they] want to believe is tremendous. And it can be quite corrosive, quite malign. It affects how the rest of us think about the origins of life, about the nature of reality ultimate questions and about how to treat the most vulnerable members of humankind, the unborn in the womb, whether with care or savage disregard.

These are reasons for applying special scrutiny and an extra degree of skepticism they are reasons for thinking your own thoughts rather than be spoon-fed when considering what Scientists Say.

For more on the culture of contemporary science, see Egnors post, Jeffrey Epstein and the Silence of the Scientists.

Photo: A baby in the womb at 17 weeks, by Nogwater, via Flickr (cropped).

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Scientific Decadence and the Myth of Objectivity - Discovery Institute

Carly Pearce’s Next Album Will Showcase Her ‘Evolution’ – kezj.com

Carly Pearce's career and life have changed substantially since she put out her full-length studio debut, 2017'sEvery Little Thing. In that time, she's earned her first No. 1 hit, scored major awards and nominations, toured all around the world, gotten married and come into her own as an artist and performer. Now, with the release of her self-titled sophomore album just around the corner, Pearce says she can't wait to catch fans up on all the changes in her life.

"So much has happened to me that has made me, I guess. That first record, I was searching and hoping that people in country music would embrace me. Now I know that I have a place in the genre, which is amazing," Pearceexplained at the 2019 ASCAP Country Music Awards last November.

"I had, just, more confidence with that [when I was making my second album]. So many great things happened to me that gave me the confidence to know, 'People want to hear what you have to say,'" she continues.

One of Pearce's biggest champions as she worked onCarly Pearcewas her producer,Busbee, who helped her develop her musical sound and incorporate her personal life into her next batch of songs. "I told Busbee when we went in, I wanted it to be an evolution of music as a woman," Pearce says.

In late September, Busbee died at the age of 43, following a battle with brain cancer. In light of that loss, Pearce says her new project has taken on an even greater meaning.

"Obviously, it's taken on a little bit of a bittersweet thing for me, because it was the last record that [Busbee] finished before he passed away, tragically," Pearce notes. "That takes on an extra special meaning for me. I feel like it's my duty to make him proud with this music."

Now that the album's release is less than a month away, though, Pearce says her overarching feeling is excitement at being able to finally let fans in on a new chapter of her life. "[The record] has been finished for a while, so just being able to finally have an end date to it, and to be able to put these songs out, [is amazing]," she adds.

Who Else Is Releasing New Music in 2020?

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Carly Pearce's Next Album Will Showcase Her 'Evolution' - kezj.com

Temtem Guide – How to Evolve your Temtem – Attack of the Fanboy

If you want to fill out your collection of Temtem and gather data on all the multitude of species available to catch and battle in the game, you need to understand evolution. This mechanic, adopted from the franchises and series that inspired Crema to create the newly released MMORPG, is key to becoming the best player you can, with the strongest team of Temtem at your disposal. But as a new franchise, players are wondering how everything works. So heres how to evolve your Temtem.

First, a warning. This is a bit too early for all the details of evolving your Temtem as the game is still in Early Access and a lot of this can change. Also this means that currently were still discovering elements of the game and its mechanics, so be sure to let us know if we get anything wrong. With that said, well share as much as we can and keep this article updated as the game works its way to full release in 2021. So, how do you evolve Temtem? All the ones we currently know about use the tried and true method of certain level caps.

Most creatures in Temtem need to be leveled up a certain amount of times, in most cases seven levels, before they will evolve. For example Crystle, one of the starter Temtem, will evolve into Sherald at level 30. The more you use it in battle, the quicker youll hit this level, and the sooner it will evolve. However, this isnt the only thing to keep in mind in regards to evolution in this new game.

Many Temtem follow the usual formula of hitting a certain level then evolving into Stage 2 or possibly Stage 3 versions of that species. Some Temtem lack further stages though and will never evolve, so watch for more on that as we dig into the game. However others have even more complexity to them such as Meta-mimetic Tetem which can evolve into different final forms depending on certain conditions. Well break these down once theyre fully discovered and fleshed out.

So thats what we know now about how to evolve your Temtem. Theres lots more to discover as the game gets updated and the deeper mechanics are added in. Be sure to check back for more as this develops.

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Made in the Tri-States: For longtime Dubuque manufacturer, evolution a key to longevity – telegraphherald.com

Given its extensive history in Dubuque, one might assume The Adams Co. has been a model of consistency for the past 130 years.

During that span, however, the companys path to success has been marked by frequent and significant changes. Adams Co. moved its location across town earlier this century and, at various points, overhauled its business model.

Company executives speak about this change with a sense of pride.

We have evolved substantially from what our roots were, said President and CEO Steve Arthur.

Today Adams Co. is located 8040 Chavenelle Road in Dubuque. The business employs 70 people and custom-manufactures gears, shafts and power transmission parts.

The companys history dates back to 1883, when it operated primarily as a foundry out of a facility near Dubuques Ice Harbor. It continued to serve in that capacity for more than a half-century.

Other areas of focus have come and gone.

For the better part of 20 years, spanning the late 1800s into the early 1910s, the company also was in the business of car manufacturing. It produced multiple vehicles under the name Adams-Farwell but ultimately left that line of work when the likes of Henry Ford carved out a massive market share in the industry.

Adams Co. also created decorative fireplace accessories for about three decades until selling off that portion of the business in 2008.

While its focus may be narrower today, the companys methods continue to evolve and leaders are continuously looking for ways to improve in an ever-changing market.

Adams Co. creates products for a wide range of clients spanning multiple industries and geographical areas.

The business primarily creates gears and shafts for customers in the ag, oil and construction industries. Its reach extends well beyond its Midwest roots.

We have customers in nearly every state, said Pat Meehan, a sales coordinator who also works in the tooling and methods department.

Meehan noted that Adams Co. has some customers in Canada and one in Europe. Meanwhile, many of its domestic customers disperse parts from the Adams Co. across the globe, meaning that the Dubuque-based firm has more of an international reach than meets the eye.

Generally speaking, Meehan said most products created at Adams Co. go through a similar process.

The company receives steel in large bars, measuring from 20 to 30 feet in length. Employees then saw the steel into the preferred lengths.

From there, the parts are taken to the blanking department where they are transformed into the general configuration that is desired.

Parts are then sent to the tooth-cutting department, where the teeth of the gears are etched into the product. Afterward, the products get burred, a process that involves removing the sharp edges.

The parts are ultimately sent to an outside facility where they undergo heat treatment. This process ensures a hardened and more durable surface.

While many products follow a general path to completion, Meehan emphasized that each part is tailored specifically to a clients needs.

Everything we make is per our customers designs, Meehan said.

For more than 120 years, Adams Co. called the Ice Harbor home.

In the early 2000s, however, efforts to reshape the Port of Dubuque into a more tourist-friendly location precipitated an end to this lengthy run. Officials from the City of Dubuque and Adams Co. worked in conjunction on a relocation plan that ultimately prompted the companys move to Dubuque Industrial Center West.

In 2004, the company moved into a newly constructed, 50,000 square-foot building that sits on 9 acres along Chavenelle Road.

The company now has resided in that same facility for 15 years. However, the equipment and processes within continue to change rapidly.

With what we do, there is a constant evolution in the types of tooling being made, the material they are made out of and the coatings on them, said Meehan. It affects the type of equipment we invest it, how we make the parts and how quickly things get done.

Arthur said Adams Co. now employs about 70 workers and said that number has remained relatively stable in recent years. The productivity of the company has continued to rise while the workforce total remains level.

Employees are more efficient because the processes are more efficient, said Arthur. Weve been able to increase our levels of work without increasing the workforce.

Like many American manufacturers, Adams Co. continues to be impacted by foreign competition.

Arthur acknowledged that the company is acutely aware that it is operating in an international market. As a result, it will continue to emphasize innovation.

We are always looking to apply our knowledge to other types of industries and product lines, he said. As the world changes, you have to change with it. We will continue to try to evolve.

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Made in the Tri-States: For longtime Dubuque manufacturer, evolution a key to longevity - telegraphherald.com

Shrinking dinosaurs and the evolution of endothermy in birds – Science Advances

Abstract

The evolution of endothermy represents a major transition in vertebrate history, yet how and why endothermy evolved in birds and mammals remains controversial. Here, we combine a heat transfer model with theropod body size data to reconstruct the evolution of metabolic rates along the bird stem lineage. Results suggest that a reduction in size constitutes the path of least resistance for endothermy to evolve, maximizing thermal niche expansion while obviating the costs of elevated energy requirements. In this scenario, metabolism would have increased with the miniaturization observed in the Early-Middle Jurassic (~180 to 170 million years ago), resulting in a gradient of metabolic levels in the theropod phylogeny. Whereas basal theropods would exhibit lower metabolic rates, more recent nonavian lineages were likely decent thermoregulators with elevated metabolism. These analyses provide a tentative temporal sequence of the key evolutionary transitions that resulted in the emergence of small, endothermic, feathered flying dinosaurs.

The evolution of endothermy in birds and mammals is regarded as one of the most important transitions in vertebrate evolution, providing an extraordinary case of evolutionary convergence between these groups that was pivotal to their widespread geographic distribution and ecological success (1). While several groups of invertebrates and vertebrates can raise their temperatures above ambient, the maintenance of high and constant body temperature (Tb) through endogenous heat production at rest is exclusive to birds and mammals and explains their greater mobility, stamina, and tolerance to a wider range of conditions. Nonetheless, because this strategy is energetically costly and leaves virtually no trace in the fossil record, the tempo and mode of the evolution of endothermy remains one of the most contentious subjects in vertebrate evolution (26). To understand how, when, and why endothermy arose during the evolution of birds and mammals, two fundamental questions must be considered: What are the costs and benefits of this strategy when compared against ectothermy, and, more importantly, under which conditions would the transition toward endothermy be favored?

Here, we address these questions using the Scholander-Irving model of heat transfer (Fig. 1) (7, 8). For any organism in thermal steady state, the model states thatMR=C(TbTa)(1)where MR corresponds to metabolic rate (milliliters of O2 per hour), C corresponds to thermal conductance (milliliters of O2 per hour and degrees Celsius), and Tb Ta constitutes the thermal gradient between body and ambient temperature (degrees Celsius), respectively. While this relationship has been used to study thermoregulation in endotherms for more than 60 years (79), it has been rarely used for ectotherms because of their low MR and high C that results, as the ratio MR/C tends to zero, in the rearranged approximation Tb = Ta [but see (10, 11)]. However, because all living organisms produce endogenous heat, the model remains applicable under thermal steady state, which is a crucial assumption to circumvent the use of complex dynamic models often applied to ectotherms that would render analyses below intractable.

(A) The cost-benefit to switch from ectothermy to endothermy for different ranges of body size was quantified with the Scholander-Irving model, which describes how a rise in metabolism at rest (cost) increases the thermal niche Tb Ta (benefit). Because there is no thermal gradient between the organism and the environment in the absence of heat production, this curve intersects the abscissa at Tb = Ta when MR = 0 (8). The solid blue and red lines depict the metabolic curves of a typical ectotherm and endotherm, respectively, and the open symbols depict the maximal thermal gradient Tb Ta possible with resting metabolic rates, used in our model (Eq. 2). (B) A reduction in body size, consistent with the one described from ancestral theropods to basal birds (22), constitutes the evolutionary path of least resistance as the energy costs of being large are traded for those of being endothermic.

The costs of endothermy can be quantified as mass-independent energy expenditure, whereas the benefits include greater mobility and foraging efficiency, predator avoidance, tolerance to, and colonization of a wider range of environmental conditions, increased growth rates, and homeostasis (1). Many of these benefits derive from an elevated Tb, which can only be maintained above a certain minimum Ta. Therefore, Tb Ta quantifies the thermal niche that organisms can occupy, and its expansion can be used to estimate the net benefit of endothermy (Fig. 1). The cost-benefit of adopting an endothermic lifestyle may now be calculated as the fold increase in MR required to expand the thermal niche by 1C (hereafter cost per degree), which can be worked out for a constant Tb ascost per degree=(MRendo/MRecto)(TaendoTaecto)(2)with the subscripts referring to the ectothermic ancestor and the endothermic descendant (Fig. 1). This index of energy cost per degree Celsius is expected to change with body size because both MR and C vary allometrically (Fig. 2) (12, 13). For example, since a generic endotherm exhibits a 5-fold higher MR adjusted to Tb = 38C (14) and a 2.5-fold lower C than an ectotherm (11) (allometric equations in Fig. 2), then the size reduction from the estimated ~370 kg for the basal Tetanurae to ~0.9 kg for the basal bird would result in reduction in total energy expenditure from 10,194.0 to 574.4 ml O2/hour and a thermal niche expansion of 12.1C (from a thermal gradient of 6.7 to 18.8C) (Fig. 2). This corresponds to a cost per degree of 4.65 103 per C, or 7.1 and 1.6% of the predicted costs should endothermy have evolved in lineages with a constant size of 370 and 0.9 kg (cost per degree of 6.48 102 and 2.89 101 per C, respectively) (Fig. 3). These calculations, which can be replicated with the exact body size estimates provided in the Supplementary Materials (see Methods), show that the evolution of smaller sizes reduces the energy costs to evolve endothermy, as originally proposed by McNab for mammals (3). In the next sections, we explore how this heat transfer model, combined with well-resolved phylogenies and body size reconstructions, can shed light on how endothermy evolved in birds from their theropod ancestors.

(A) Theropod phylogeny (15) with branches color-coded according to reconstructed metabolic levels. (B) Scaling of metabolic rate versus body mass (12) for ectotherms (MR = 0.68mass0.75) and endotherms (MR = 3.4mass0.75) and the predicted trajectory of the bird stem lineage during the transition from ectothermy to endothermy. Dashed lines show fold differences between ectotherms and endotherms (1 to 5); open and closed symbols depict reconstructed values for the bird stem lineage and the tips of the phylogeny, respectively (see Methods). (C) Scaling of thermal conductance C and body mass (13) for ectotherms (C = 2.5mass0.5) and endotherms (C = 1.0mass0.5), fold differences from 2.5 to 1. (D) Thermal gradient and fold differences calculated with Eq. 1 and values in (B) and (C). The log-log linear trajectories connecting MR and C of the ectothermic ancestor and the endothermic descendant, as well as the resulting trajectory in thermal gradient, are shown with the continuous lines.

(A) The miniaturization from Tetanurae to basal birds inferred from the fossil record (15), contrasted against 100 simulated size trajectories starting from the same ancestral body size for illustrative purposes (note that for the subsequent full null model, the ancestral body size is allowed to vary). The error represents the SD in reconstructed values across 20 candidate trees (see Methods). (B) The frequency distribution of body mass ratios obtained across 10,000 simulated body size trajectories (histogram) and the energy costs to evolve endothermy expressed per degree Celsius (Eq. 2) under this null model (gray symbols). In this case, the ancestral body size was obtained from a uniform distribution ranging between 10 g and 100,000 kg. The empirical estimate in the bird stem lineage is shown in red. The region in which a reduction in body size would compensate for the energy costs of evolving endothermy, enabling the population to increase in a scenario of constant resources, is highlighted in gray. The arrow depicts the expected population fold increase, given the observed body size reduction in the bird stem lineage as endothermy evolved. These analyses indicate that the energy costs to evolve endothermy are reduced with miniaturization and, as a result, population size may have increased despite the metabolic costs of an endothermic lifestyle.

We estimated the costs of evolving endothermy along the bird stem lineage from reconstructed ancestral body sizes inferred from the fossil record (15). To quantify the energy costs in alternative scenarios, we simulated the evolution of body size along this lineage and obtained the distribution of cost per degree under this null model (Fig. 3). We assumed an undirected Ornstein-Uhlenbeck (OU) model of size evolution bounded between 10 g and 100,000 kg and a mean evolutionary rate equivalent to values reported for these theropods (see Methods). Estimates of cost per degree across different simulated size trajectories were log-normally distributed with a fold-change median of 0.124 per C (100.9 in Fig. 3B), which implies that, relative to the metabolism of the ectothermic ancestor MRecto (Eq. 2), the energy cost to increase the thermal niche by 1C as endothermy evolves amounts to roughly 12.4% at a constant body size. According to our null model, 95% of simulated costs per degree Celsius would fall between 275 and 0.576% depending on whether body size increases or decreases, respectively. In this context, simulations indicate that the energy costs per degree Celsius decrease markedly with miniaturization (Fig. 3). Two phenomena explain these reduced costs. First, the expansion in thermal niche following an increase in MR is disproportionally higher in larger ectotherms because they can maintain a high Tb with a relatively low mass-independent MR due to inertial homeothermy (also known as gigantothermy) (3, 1618). Accordingly, the residuals of cost per degree controlling for the fold difference in size between ancestor and descendant are negatively related with the simulated ancestral size according to a linear regression (F1,9998 = 1.94 105, r2 = 0.95, P < 0.001). That is, the larger the starting size of the ectothermic ancestor, the cheaper the transition to endothermy should be. Second, during miniaturization, the energy costs of being large are traded for being endothermic, which helps to explain how high energy turnover rates evolved despite their impact on food and water requirements. Birds require between 15 and 20 times more food than a similar-sized reptile (6), which could be problematic because the proportional fold reduction in population size expected if resources were constant might jeopardize the populations long-term persistence in evolutionary time (19, 20) (certainly, the benefits of being endothermic and capable of obtaining more resources might partly offset this limitation).

In contrast, in our model, energy equivalence between an ectotherm and an endotherm is attained with an 8.55-fold decrease in body size (Fig. 3); thus, a 43.3-kg bird should have the same requirements as its 370-kg Tetanurae ancestor, everything else being equal [life is certainly more complicated, and differences in activity patterns or home range size between ectotherms and endotherms should affect this rough estimate (21)]. Assuming that energy can be assigned to either body size or abundance, our analysis shows that, irrespective of the potential increment in food resources resulting from an expanded thermal niche or higher access to small prey as size decreased, populations could still exhibit a 30.5-fold increase as endothermy evolved (Fig. 3). This might explain how, despite the inherent variation in resource availability expected in evolutionary time, smaller sizes and higher energy turnover rates may have been systematically favored in this lineage. Accordingly, the estimated costs of 0.466% per C estimated for the bird stem lineage are significantly lower than our null distribution (one-tailed P = 0.0172; Fig. 3) and, therefore, energetically cheaper than most simulated scenarios using realistic background rates of body size evolution for theropods. Results remained qualitatively identical for other null models with relaxed assumptions such as a smaller Tetanurae ancestor or assuming Brownian motion model of evolution (see Methods).

The size reduction in the bird stem lineage (15, 22, 23) closely matches the theoretical path of least resistance for endothermy to evolve. We now reconstruct how this phenomenon might have unfolded in the theropod phylogeny. Combining node dates and body size estimates (15) with the allometric shift that would describe the transition to endothermy, we interpolated MR and C of intermediate ancestors in this lineage (Fig. 2). This procedure indicates that the rise in MR spanned most of the Early-Middle Jurassic [~180 to 170 million years (Ma) ago] (Fig. 4) and involved theropod groups in which the occurrence of protofeathers and feathers was already ubiquitous (24). It also suggests that metabolic levels were highly diverse across contemporaneous lineages of Coelurosauria, Maniraptora, and Paraves, which might partly account for the emergence and diversification of these groups during the Late Jurassic (22, 25), and the abnormally high diversity of Coelurosauria at intermediate body sizes (between 30 and 300 kg) when compared against other dinosaur groups (15). A niche-filling model of adaptive radiation in Mesozoic dinosaurs also detected exceptional rates of body size reduction in the bird stem lineage, particularly in the basal nodes of Coelurosauria and Paraves, although no suitable evolutionary hypothesis was proposed to account for this result [table 3 in (23)]. Our analyses show that the evolution of an endothermic machinery, concomitantly with the resulting thermal niche expansion, provides a plausible explanation for both the radiation and the reduction in size detected in these lineages.

(A) Reconstructed temporal course of metabolic evolution in the bird stem lineage, with dashed lines showing how reconstructions change assuming that either Paraves or Neornithes were fully endothermic instead of the basal bird [for calculations with Neornithes, we assumed a body size of 150 g based on estimates for Vegavis (22) and a time estimate of 100 Ma ago (39)]. The fold increase in MR was calculated by dividing the reconstructed MR during the transition to endothermy by the MR expected for a similar-sized ectotherm and is therefore dimensionless and independent of body size. (B) The evolutionary path of least resistance from ectothermy to endothermy includes inertial homeothermy as a transitional stage, followed by an increase in metabolism concomitantly with a reduction in size. (C) Hypothetical sequence of evolutionary transitions in the bird stem lineage, which combines results from this study with phylogenetic reconstructions of epidermal structures (24, 42) and capacity for active flight (38) (see the main text).

Two exceptional phenomena are observed during the evolution of birds: a sustained (but not necessarily gradual) miniaturization lasting millions of years and the emergence of endothermy. We argue that these phenomena are mechanistically linked. Our reconstruction suggests that endothermy evolved concomitantly with the decrease in size along the bird stem lineage, as originally proposed for mammals by McNab (3), and that related theropod clades should exhibit a whole spectrum of MR. Although it may be debatable to what extent energy costs are minimized in evolution, such a principle has been widely invoked or implicitly assumed to explain the diversity of thermoregulatory strategies across extant lineages (79) and may be equally useful to study how such a diversity evolved. The proposed scenario explains the conundrum of an expensive lifestyle being systematically favored despite its energy costs and explains the sustained miniaturization that preceded the origin of birds (15, 22, 23, 26), the so-called mesothermy (12), and intermediate to high growth rates of many dinosaurs (12, 2732). Previously labeled mesotherms (12) are either inertial homeotherms such as tunas, leatherback sea turtles, and large dinosaurs or small endotherms such as echidnas and, according to our reconstruction, the maniraptor Troodon and Archaeopteryx (Fig. 2). In this context, mesothermy constitutes an ambiguous concept from a mechanistic perspective because elevated MR due to thermodynamic effects (i.e., a high Tb due to thermal inertia and a large body size) is confounded with high MR due to the evolution of increased mass-independent energy turnover rates and true endothermy.

Inertial homeothermy might, in fact, constitute a necessary transitional state (3, 1618) in which homeothermy and a high thermal gradient Tb Ta can be maintained at low metabolic costs. That is, we posit that the MR of the large ancestral theropods (>300 kg) would fall in the allometric curve for ectotherms (21) and, yet, these organisms would be able to maintain a thermal gradient more in line with that of extant endotherms (Fig. 4B). Subsequently, selection toward smaller sizes would favor elevated MR if these large ancestral theropods were physiologically committed to homeothermy, as discussed by McNab (3), which would explain the departure from the ectothermic metabolic allometry with miniaturization (Fig. 2A). While this proposition has been generally dismissed (6, 33, 34) on the basis that a large size obviates thermoregulatory needs for high metabolic rates (16), this counterargument is true only within a limited range of Ta and body sizes and neglects other ecological advantages of a high mass-independent aerobic capacity and the advantage to a high reproductive rate, growth rate, etc. In large terrestrial animals, selection on increased MR for sustained activity, parental care, or high growth rates, presumed drivers of the evolution of endothermy in birds and mammals according to different hypotheses (4, 5), should inevitably increase the thermal gradient Tb Ta and give rise to larger thermal niches as a useful by-product (Fig. 2). In evolutionary time, it is only reasonable to expect lineages to exploit newly opened niches and eventually diversify (35). Under this scenario, thermoregulatory performance is expected to evolve regardless of the selective pressures favoring a high aerobic capacity, which reconciles alternative theories on the evolution of endothermy in birds and mammals (26).

While quantitative estimates are expected to vary with the allometric relations used, these patterns should be robust to variation in scaling and violations of the models assumptions. On the basis of Eq. 1, as long as the MR scaling exponent remains greater than that of C, the thermal gradient Tb Ta should increase with size and energy costs should decrease. In addition, the greater empirical MR scaling exponent described for ectotherms (12, 14) would result in lower energy costs per degree Celsius at larger sizes, whereas the lower Tb of smaller ectotherms should disproportionally increase the energy requirements to attain endothermy within this size range. The thermodynamic constraint imposed by a lower Tb would also buffer any potential advantage of a higher aerobic capacity on performance (i.e., highly aerobic lizards remain inactive when they are cold); hence, some degree of homeothermy would be desirable for high MR to evolve. In this context, it is important to recall that our model assumes a constant Tb for practical reasons, whereas, in reality, Tb likely varied across groups and was possibly higher and more stable in larger lineages, everything else being equal, due to a reduced surface relative to volume (36). Inherently higher and stable Tb in larger dinosaurs might partly explain their intermediate metabolic levels between reptiles and extant mammals and birds and may have contributed to the evolution of endothermy by facilitating parental care and favoring higher growth rates in these lineages (32).

Admittedly, the size reduction immediately preceding the radiation of birds could also be related with the evolution of flight, and it is quite possible that paravians or earlier groups were fully endothermic. The intermediate growth rates of these groups suggest otherwise (12, 2732), and given the accelerated rates of body size reduction in this period, this possibility does not alter the general trend of MR evolution reported here (Fig. 4). Moreover, relative forelimb elongation and increased flapping assisted locomotion are detected primarily within paravians (37, 38). Consequently, the stepwise evolution in body size reported by Benson et al. (15), with a first sustained reduction between the ancestors of Tetanurae and Paraves and a second shift within Avialae roughly 20 Ma later, might be associated with, respectively, the evolution of endothermy and flight. Alternatively, if basal birds were not fully endothermic and this derived condition emerged later in groups such as crown group birds (Neornithes) (39), as suggested by the slow growth rates of Archaeopteryx (27), then these traits may have coevolved in tandem, to some extent, within the avian lineage. Flightless birds exhibit lower MR (40), and there is substantial variation in metabolic levels among extant birds (41); hence, these alternatives are not mutually exclusive. On the contrary, one should perhaps envision a scenario in which aerobic capacity shifted multiple times along the theropod and avian phylogeny (Fig. 2), resulting in a gradual rise in MR above the allometry of extant reptiles and a whole gradient of metabolic levels in these groups.

In any case, comparative analyses suggest that feathers or protofeathers evolved before the emergence of Coelurosauria (24, 42) and the rise in MR reconstructed in our study (Fig. 4). Together, the available evidence indicates that the evolution of flight cannot explain the reduction in size around the Early-Middle Jurassic boundary interval (~180 to 170 Ma ago). More crucially, our results, combined with previous analyses on the evolution of body size, feathers, and flight in the bird stem lineage (15, 2224, 26, 37, 38, 42), give rise to a relatively well-defined temporal sequence of key evolutionary transitions and a detailed working hypothesis for future studies (Fig. 4). This interpretation, that endothermy preceded the evolution of flight, is also consistent with descriptions of skeletal pneumaticity in derived, but not basal theropods (43), and mirrors the evolutionary sequence that can be inferred for bats since endothermy is a plesiomorphy present in virtually all mammals. Whether a similar scenario could explain the evolution of flight in pterosaurs, which have been recently described to exhibit feather-like integumentary structures (44), remains an open question. The elevated MR among bats and the enormous variation in metabolism across mammalian groups constitute a reminder that endothermy (and ectothermy, for that matter) constitutes a matter of degree rather than kind, which might explain why the earliest dinosaurs may have exhibited higher metabolic rates than those of extant reptiles (45) and early bird-like taxa growth rates that were not quite comparable to modern birds according to bone histology (27, 32). Thus, these findings are not necessarily at odds with our proposition that the marked reduction in body size during the evolution of the bird stem lineage was accompanied by a major shift in metabolic levels; they simply highlight that there is likely more to the evolution of endothermy in extinct archosaurs, dinosaurs, and birds than our analyses can convey.

To quantify how the sustained reduction in body size along the avian lineage would affect energy expenditure during the evolution of endothermy, we used the theropod phylogeny and body mass estimates reconstructed from the fossil record reported by Benson et al. (23) to build a realistic null model. We cross-validated this dataset against an independent study (22), which includes other taxa and is based on different methods to estimate body mass, phylogenetic relations, and evolutionary trends. For the species shared between these studies (n = 94), body mass estimates were very similar according to a regular regression (slope, 1.10 0.02 SE; r2 = 0.98, P < 0.001), and so was the structure of the phylogenies in both studies based on Bakers , which estimates the similarity between two trees of hierarchical clustering and varies between 1 and 1 as a regular correlation ( = 0.962, P < 0.001) (figs. S1 and S2). While both studies reach very similar conclusions and detect exceptional body size reduction in nodes along the bird stem lineage (22, 23), there are two remarkable differences: Lee et al. (22) reported a sustained, gradual decrease in body size along the bird stem lineage from a large ancestral theropod (175 kg), whereas Benson et al. (23) suggested that this miniaturization occurred in a stepwise fashion and later in time, following a period of size increase in early theropod evolution from a smaller ancestor (10 to 30 kg). Because the larger ancestral size seems to be an artifact of incomplete and biased sampling of early taxa (15) and the stepwise decrease in size was observed by Novas et al. (46), estimates used here correspond to unweighted averages of reconstructed ancestral body mass and divergence times obtained across 20 candidate phylogenies by Benson et al. [appendix S5 in (15)]. The dataset used does not change the general conclusions of this study, and analyses using body mass data and phylogeny of Lee et al. (22) are shown in the Supplementary Materials (figs. S3 to S5).

Allometric equations for endotherms were obtained from the literature for MR (12) and C (13). For mathematical tractability and to ensure that results were comparable across different ranges of body size, we obtained parallel curves for ectotherms by dividing the intercept of these equations by 5 (14) and 2.5 (11). On the basis of these allometric curves, we then assigned typical ectothermic and endothermic values for the Tetanurae node (~370 kg, range 290 to 420 kg) that corresponds to the largest theropod ancestor in most reconstructions (15, 22) and for the basal bird (~0.93 kg, range 0.78 to 1.10 kg). Results are expressed as fold change in MR or C with respect to the ectothermic allometric curve throughout the study, calculated as the ratio of the observed estimate/expectation for a similar-sized ectotherm. Subsequently, from linear log-log MR and C curves connecting these two taxa (Fig. 2, B and C), we interpolated how these variables evolved in the bird stem lineage using the body size estimates reconstructed for intermediate nodes. With the MR and C calculated for these nodes combined with divergence time estimates, we then reconstructed how metabolic levels evolved along the bird stem lineage. This approach assumes, for simplicity, a constant rate of fold change in MR and C as body size decreases, which is unlikely. Nonetheless, the exceedingly high rates of body size reduction observed primarily between Neotetanurae and Paraves (15, 22, 23) constrain the period in which metabolic levels increased in a narrow temporal window, which should remain largely unaffected by the general shape of the evolutionary path connecting the end points (i.e., the ectothermic ancestor and its and endothermic descendant). For the remaining species in the theropod phylogeny, we assumed that they inherited the fold change in MR and C from their most recent ancestor in the bird stem lineage, shifted the intercept of the allometric curves accordingly to obtain appropriate estimates of MR and C accounting for size effects, and then calculated Tb Ta with Eq. 1.

After quantifying the energy costs of evolving endothermy for the basal stem lineage leading to birds (Eq. 2), we built a null distribution of energy costs by simulating 10,000 different body size trajectories along this lineage under a null model. The body size distribution for the ancestral ectotherms was sampled from a uniform distribution ranging between 10 g and 100,000 kg, which encapsulate the range of body sizes observed within the theropod phylogeny, and the distribution of their descendent endotherms was constrained within this same range by removing those replicates falling outside this range. These boundaries are included to ensure that the body size null distributions fall within the range observed across higher vertebrates, which presumably reflect biomechanical or physiological constraints (e.g., simulating a 1-g endotherm is not realistic as sustaining a thermal gradient at this size is not really possible).

Body size evolution was simulated with an OU process, using a code written ad hoc. We used the 2 and parameters fitted to 20 candidate theropod phylogenies by Benson et al. (15), which measure, respectively, the intensity of random fluctuations in the evolutionary process and the strength of selection toward a presumed optimal trait value . To be as conservative as possible, from the seven single-optimum OU models with the best fit reported by Benson et al. (15), we used the parameters that resulted in the highest phenotypic variance following 10,000 diagnostic simulations: 2 = 0.025 and = 0.005 (obtained from the Theropoda tree 1 in their appendix S5; see figs. S6 and S7 for additional details on the null model and selected parameters). In addition, to remove the contribution of directional trends in these models and obtain random variation comparable to the empirical data, we used the same and 2 fitted for each tree and setting to the ancestral body mass [appendix S5 in (15)]. The amount of time between the ectothermic ancestor and endothermic descendant was set to 47 Ma, or the average difference (46.6 2.9 Ma) between the Tetanurae and basal bird nodes across the candidate phylogenies (15).

With this approach, we obtained a null distribution of energy costs to evolve endothermy from theropods to paravians under different body size trajectories, controlling for the amount of time available for this transition and using body size evolutionary rates inferred from fossil data. Subsequently, we tested whether the cost per degree estimated for the bird stem lineage was lower than that of 95% of the simulated datasets, which would indicate that the empirical trajectory is energetically cheaper than the null hypothesis holding type I error rates at 0.05, and suggest that the observed miniaturization constitutes an evolutionary path of least resistance from an energetic point of view. Results using a Brownian null model setting = 0.0, which is more conservative as the body size variance under this null model is expected to increase, were qualitatively identical (fig. S8). Similarly, results from our null model were also statistically robust to uncertainty in the ancestral body size of the Tetanurae node, whose estimated range between 290 and 420 kg always resulted in reduced energy costs associated with miniaturization with a P < 0.05.

Supplementary material for this article is available at http://advances.sciencemag.org/cgi/content/full/6/1/eaaw4486/DC1

Fig. S1. Relationship between body size reconstructions performed by Benson et al. (15, 23) and Lee et al. (22).

Fig. S2. Comparison between the topologies of the theropod phylogeny reconstructed by Lee et al. (22) and Benson et al. (15, 23).

Fig. S3. Replicate of Fig. 2, except that, in this case, analyses were replicated using the dataset and phylogeny by Lee et al. (22).

Fig. S4. Replicate of Fig. 4, except that, in this case, analyses were replicated using the dataset and phylogeny by Lee et al. (22).

Fig. S5. Comparison between reconstructed metabolic levels along the bird stem lineage using the dataset by Benson et al. (15) and Lee et al. (22), plotted against the 1:1 line.

Fig. S6. Phenotypic variance simulated with the difference parameters fitted by Benson et al. (15) for the theropod phylogeny (parameters available in their appendix S5).

Fig. S7. Simulated OU model overlapped against the empirical data from Benson et al. (15) (their appendix S5), which shows that this model can replicate the distribution of phenotypic data observed along the theropod phylogeny and provide a valid null model in the absence of directionality (see below).

Fig. S8. Results from the null model in the main text compared against expectations for a more conservative model assuming Brownian motion.

This is an open-access article distributed under the terms of the Creative Commons Attribution-NonCommercial license, which permits use, distribution, and reproduction in any medium, so long as the resultant use is not for commercial advantage and provided the original work is properly cited.

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Shrinking dinosaurs and the evolution of endothermy in birds - Science Advances

Himmelfarb and Her Haters – Discovery Institute

Editors note: Historian and Darwin skeptic Gertrude Himmelfarb died on Monday, December 30, 2019. While mourning the passing of this great scholar, we are pleased to republish Professor Flannerys 2009 essay, below. See also Flannerys tribute, Farewell to Gertrude Himmelfarb, Brutally Honest Historian of the Darwinian Revolution.

If you have no enemies, it is a sign fortune has forgot you. Thomas Fuller, Gnomologia, 1732

Noted physician Thomas Fuller was an expert on eruptive fevers, and so it seems fitting to open this essay with his wry but telling observation on enemies in public life, for perhaps no contemporary historian has spawned more eruptive fever over an analysis of the reigning secular creation myth demigod, Charles Darwin, than has the present subject of this essay. If Fuller is any judge, fortune has indeed remembered Gertrude Himmelfarb.

Such fortune appeared a few months ago when Pandas Thumbused the occasion of Irving Kristols death on September 18 to denigrate Gertrude Himmelfarbs fifty-year-oldDarwin and the Darwinian Revolutionas a terrible book . . . demonstrating a lack of understanding of biology and a warped view of Darwins influence. The article, written by Jeffrey Shallit, glibly casts aspersions on the late Kristols ethics for reviewing Gertrude Himmelfarb (aka Bea Kristol) inEncounterand failing to disclose that he was the authors husband (though this writer could find no evidence of that at least with herDarwin), this without once reflecting on the questionable propriety of turning what should have been either a respectful obituary or complete silence into an opportunity to insult both the deceased and his widow. If that isnt unethical, it is at least indecent. Shallits one-sided, high-toned moralizing aside, as the Darwin year draws to a close and given the fact that Himmelfarbs biography of Darwin itself has just marked its golden anniversary, perhaps a careful reflection upon that effort is in order. What can be said ofDarwin and the Darwinian Revolutionin the dusk of 2009, fifty year after its original publication?Isit a terrible book?

Regardless of what one may think of Gertrude Himmelfarbs work, her preeminent role as an important (albeit controversial) historian cannot be doubted (seeJewish Women: A Comprehensive Historical Encyclopedia). A prolific writer, this professor emeritus of the City University of New York has not shrunk from boldly decrying moral relativism and the so-called new history, positions for which she earned widespread praise and condemnation. Never-theless, her serious consideration for the position of Librarian of Congress in 1987 is a measure of her significance as one of Americas leading scholars and intellectuals.

Thus, it would seem worthwhile to probe a bit deeper into Dr. Himmelfarbs study of Charles Darwin. It is worth mentioning that her Darwin biography was (and obviouslyis) as controversial as its subject. Given the authors refusal to duck or dodge tough issues, her attention to modern biologys paterfamilias was bound to form an explosive catalyst easily discerned in the ensuing reviews.

Upon its publication Charles Gillispie insisted that one must deplore the interpretation of Darwin and his work that Miss Himmelfarb offers for its hostility to science.1Similarly, another reviewer dismissed it as a misrepresentation that is dubious in the extreme.2But others saw it differently. J. F. Burnet, for example, praisedDarwin and the Darwinian Revolutionas thorough and authoritative, concluding, This is an important book for all students of nineteenth-century thought.3Another reviewer called it, a scholarly book, well organized and well written, interesting to the intelligent reader whatever his special field.4

One would think these reviewers had read entirely different books, and perhaps theydid. Himmelfarbs detractors, as witnessed in Shallits mean-spirited post, say little about why her book is terrible, presuming the charge alone is sufficient to indict and convict. Interestingly, Shallit provides evidence of the authors failing in a provocative link tagged , which sends one to a posting by P.Z. Myers on December 6, 2005 titled A critique of Himmelfarbs scientific views. Now Himmelfarb has been caught! The spuriousness ofDarwin and the Darwinian Revolutionis now laid bare! Well, thats what The Pandas Thumb would have you believe anyway.

On closer inspection, however, this critique of Himmelfarbs science is nothing of the kind. In actuality it is a strange diatribe on neo-conservatism with allusions to Leo Strauss, George Will, and (Himmelfarbs husband again) Irving Kristol, none of which has anything whatsoever to do with her purportedly terrible science. Stacking up a series of allegedly damning Himmelfarb quotes, Myers then links them to creationist claims as if the mere association were sufficient condemnation. The critique concludes with a peculiar discussion of Machiavelli, which not only has nothing to do with Himmelfarbs science, but indeed nothing to do with neo-conservatism (those interested in following the perfidity of Machiavelli should examine number one of Benjamin Wikers list of10 Books That Screwed Up the World).

The only substantive example of Himmelfarbs supposed lapsed science is Myers dismissal of her charge that Darwin failed to explain the evolution of the eye by means of natural selection. Quoting from Ernst MayrsWhat Evolution Is, Myers proclaims the light-sensitive spot and the Pax 6 regulatory gene to have solved the problem. Photosensitive, eyelike organs have developed in the animal series at least 40 times, declared Salvini and Mayr in 1977, and all the steps from a light-sensitive to the elaborate eyes of vertebrates . . . are still found in the living species of various taxa. But 32 years later James Le Fanu (and many others) remain unconvinced. Le Fanu freely admits that we now know the eye to have emerged independently at least forty times in several different forms, but this has only served to make Darwins evolutionary problem more vexing:

Each different type of eye compounds Darwins difficulty further, for then it is necessary topresupposefor each a series of fortuitous numerous successive slight modifications, conferring some slight biological advantage to its possessor. It is necessary to presuppose, for, despite much effort, there is not a single empirical discovery in the past 150 years that has substantiated Darwins proposal that natural selection, taking advantage of slight successive variations, explains the puzzle of perfection epitomized by so many different types of eyewhich remains yet more puzzling than it was in 1859.5

And Michael Behe notes that the biochemical complexity of vision is not answered by a single regulatory gene, as if a single bolt could explain an automobile. Indeed where might the biochemical evidence lie? Even something as simple as a light-sensitive spot is in reality a system requiring a rhodopsin-transducin complex to interact with phosodiesterase along with many other molecular processes. Moreover, there remains no explanation of how these complex interactions could have occurred by Darwinian processes. The fossil record, as Behe has so clearly pointed out, has nothing to tell us about whether the interactions of 11-cis-retinal with rhodopsin, transducin, and phosphodiesterase could have developed step-by step.6No, the problem has not been resolved and is, in fact, worse than when Himmelfarb wrote in 1959 andmuchworse than when Darwin wrote a century before her.

The poor example of eye evolution apparently exhausts Myers evidence against Himmelfarb.The rest of his co-called critique is, in fact, an ideological rant rather than a sober investigation of her alleged scientific transgressions. Both Shallits and Myers remaining appeal seems to be anargumentum ad populumas if scientific truth was decided solely on the basis of consensus. Such arguments, of course, vindicate a noble line of scientific facts in history like phlogiston, globulism, and humoral pathology. Jeffrey Shallit and P.Z. Myers are admittedly high on passion but weak on substance.

But Shallit and Myers arent the only ones. Besides the Himmelfarb-bashers already mentioned, Ernst Mayr more than a decade after the appearance of herDarwincomplained of its abyss of ignorance and misunderstanding.7Even Cornells historian of science L. Pearce Williams appreciated Himmelfarbs grasp of the historical forces bearing upon Darwin and his times but quickly demurred at her assessment of Darwinian evolution. Calling it filled with rather serious scientific blunders, even worse than Shallit and Myers, Williams issued the indictment without giving a single instance of said blunder.8

Why is HimmelfarbsDarwin and the Darwinian Revolutionthe book Darwinists love to hate? To find out a detailed examination of the book and its incisive analysis of Darwins evolutionary theory is in order. Of course, this is abigbiography and an exhaustive account cannot be given here, but a summary investigation will make the source of the Darwinists discomfort obvious.

Darwinis divided into six books: 1) Pre-history of the Hero; 2) Emergence of the Hero; 3) Emergence of the Theory; 4) Reception of the Origin; 5) Analysis of the Theory; and 6) Darwinism. The first four books are an interesting read and provide a valuable backdrop to the treatment that follows, but Himmelfarb is weakest on Darwins early years. She completely passes over Darwins Edinburgh period where he joined the Plinian Society in November of 1826 and attended all but one of the ensuing 19 meetings until April of 1827. According to Adrian Desmond and James MooresDarwin: The Life of a Tormented Evolutionist, this was young Charles introduction to seditious science. While this is crucial in understanding the development of Darwins theory, it will not be gleaned from this book.

Also, Himmelfarb believes that Darwin was uninterested in and ill-equipped to appreciate the philosophical implications of his theory. Probably a better suggestion is that Darwin wasnt so much disinterested in philosophy as he was just a bad philosopher, or at least a very superficial one. She as much as admits Darwins anemic reading in the field: What little reading he did in philosophy was parochial in the extreme. . . . It is difficult to take seriously a discussion that had, as its most frequently cited moralist and philosopher, the historian William Lecky (p. 375).9When Darwin appended a list of moral philosophers he had relied upon in preparing hisDescent, philosophers he assured his readers they would be familiar with, Himmelfarb notes that 26 were British and that [they] are today, quite as assuredly, entirely unknown.

Nevertheless, what Himmelfarb misses in the early years she more than makes up for in the last two books devoted to an analysis of the theory and the ideologicalismthat it would turn into. Here in these two sections more than anywhere else reside the sources of anger, revilement, and consternation for the Darwinists.

For Himmelfarb Darwins presentation of his theory is most vulnerable not in his marshalling of evidence that was thin enough but rather in his means of handling it. What Darwin was doing, in effect, she observes, was creating a logic of possibility. Unlike conventional logic, where the compound of possibilities results not in a greater possibility, or probability, but in a lesser one, the logic of theOriginwas one in which possibilities were assumed to add up to probability (p. 334). Essentially, Himmelfarb accuses Darwin of making an argument from ignorance:

As possibilities were promoted into probabilities, and probabilities into certainties, so ignorance itself was raised to a position only once removed from certain knowledge. When imagination exhausted itself and Darwin could devise no hypothesis to explain away the difficulty, he resorted to the blanket assurance that we were too ignorant of the ways of nature to know why one event occurred rather than another, and hence ignorant of the explanation that would reconcile the facts to his theory. When one botanist argued that his theory was contradicted by the fact that some forms remained unaltered through long periods of time and wide expanse of space, Darwin admitted the objection to be formidable in appearance, and to a certain extent in reality. But this did not deter him:

Does not the difficulty rest much on our silently assuming that we know more than we do? . . . . Certainlya prioriwe might have anticipated that all the plants anciently introduced into Australia would have undergone some modification; but the fact that they have not been modified does not seem to me a difficulty of weight enough to shake a belief grounded on other arguments.

Somehow the fact that no adequate explanation suggested itself today seemed a warrant for the belief that such an explanation would suggest itself in the future, and the explanation, moreover, would be bound to vindicate his theory. Thus the argument from ignorance was made the prelude to a confident affirmation:

We are far too ignorant, in almost every case, to be enabled to assert that any part or organ is so unimportant for the welfare of a species that modifications in its structure could not have been slowly accumulated by means of natural selection. But we may confidently believe . . .

It may be objected, however, that in the logic of science, as in the logic of grammar, three negatives do not normally constitute a positive.

To be sure, a scientific theory that explains equally well a variety of contradictory phenomena may still be true; there are reputable theories that cannot, in this sense, be falsified, and hypothetical reasoning is a legitimate, even necessary, scientific technique. The difficulty with natural selection, however, is that if it explains too much, it also explains too little, and that the more questionable of its hypotheses lie at the heart of its thesis. Posing as a massive deduction from the evidence, it ends up as an ingenious argument from ignorance (pp. 335-336).

This kind of writing infuriates Darwins defenders who are not used to such frank talk coming from the likes of a historian. And yet Himmelfarb is not the only one. Many have followed her in questioning Darwins logic and his argument (see, for example, Jacques Barzun,Darwin, Marx, Wagner: Critique of a Heritage,1941; William Irwin Thompson,At the Edge ofHistory, 1971; Robert Henry Peter, Tautology in Evolution and Ecology,American Naturalist, 1976, and Predictable Problems with Tautology in Evolution and Ecology,American Naturalist, 1978; Michael Denton,Evolution: A Theory in Crisis, 1985; Thomas Nagel,The View From Nowhere, 1986; Stanley L. Jaki,The Savior of Science, 1988;R. F. Baum,Doctors of Modernity: Darwin, Marx & Freud, 1988; Phillip Johnson,Darwin on Trial, 1993; David Stove,Darwinian Fairy Tales, 1995; Didier Maleuvre, Can We Believe in Darwin?,Comparative Literature, 2001; James Le Fanu,Why Us?, 2009). Nevertheless, despite skeptics, Darwins theory was able to rise as the reigning paradigm in biology and moreover retain that status to the present day. How so?

Himmelfarb offers a couple of reasons, neither of which was based upon the weight of any purported evidence. Quoting a letter from Darwin to Asa Gray on December 21, 1859,Origins author recommended that the book, released that previous month, be read by intelligent men, accustomed to scientific argument, though [curiously enough]notnaturalists. But as Himmelfarb notes, What he [Darwin] did not properly appreciate, however, was that it was less as intelligent men accustomed to scientific argument that they judged and approved theOriginthan as intelligent men susceptible to philosophical prejudice (p. 296). More specifically, it was the anfractuous nature of the argument itself that worked mischievously on its behalf:

It was probably less the weight of the facts than the weight of the argument that was impressive. The reasoning was so subtle and complex as to flatter and disarm all but the most wary intelligence. Only upon close inspection do the faults of the theory emerge. And this close inspection, by the nature of the case, was largely vouchsafed. The points were so intricately argued that to follow them at all required considerable patience and concentration an expenditure of effort which was itself conducive to acquiescence. Only those determined in advance to be hostile were likely to maintain a vigilant and hence critical attitude. In his rapid volley of expectations, where one might fail, another would hit the mark, and where one line of defense had to be abandoned, another was hastily erected. And there were few to point out that in the strategy of reason, as in the strategy of warfare, the cause was not better served by a succession of feeble defenses than by a single strong one.

More important, however, than any assets which Darwins theory might be thought to possess was the bankruptcy of his opponents. The only serious rival, as a general theory, was creation (pp. 350-351).

Himmelfarb is right. William Paleys venerable argument that a watch found in a field suggested a watchmaker was powerful analogous argument for design and teleology in nature. However, at the time he made it in 1802 few details were known about complexity in the natural world. Paleys apologetic could in many ways be answered by prior skeptics like David Hume. Paleys waxing eloquent on the divine beauty of his garden looked more like romantic effusion than sober analysis, and he often saw Gods hand in virtually everything in nature raising the question (as Darwin himself did), if God is omniscient and benevolent then whither evil and pain? Paley and the authors of theBridgewater Treatisesinsisted that all creation demonstrated Gods manifest infinite wisdom and power and goodness. Cornelius G. Hunter is quite correct in hisDarwins Proof when he notes, Natural theology was lopsided. Yes, the world is amazing, but the natural theologians happy view of nature could hardly be justified in light of the real world. It is easy to see why this version of natural theology supplied a ready source of material for its opponents (p. 90).

But there is yet a third reason why DarwinsOriginwas so readily accepted; Himmelfarb has already alluded to it when she suggested that Darwins supporters were found among those already philosophically inclined to accept it. This is an important point and one that she concludes with in assessing the nature of the so-called Darwinian revolution. Skeptic and freethinking philosophers had clearly prepared the way for a wholly naturalistic account of creation and biological life.

But what of the theory itself? Once launched, how did Darwin himself handle the responses to and further development of his evolutionary theory? Here Himmelfarb is at her best. In order to fully appreciate Darwins theory as it blossomed (perhaps metastasized is a better word) theDescent of Manmust also be examined. It is clear that, over time, Darwin demonstrated a discernable retreat from his theory of natural selection, instead turning to two subsidiary but increasingly important notions to address assorted problems: one was pangenesis, the other was sexual selection. Pangenesis proposed that gemmules, shed by body cells and containing hereditary information, collect in the reproductive organs and play a key role in inheritance. For Darwin, pangenesis (today thoroughly discredited) explained blended inheritance, reversion to ancestral types, limb regeneration, and even Lamarckian concepts of use and disuse. Of course genetics changed all of this. There is no inheritance of acquired characteristic.When Dutch botanist Hugo De Vries tried to create a permanent change of type by the selection of existent variations, he found that he could not. When his ear of corn with an extra row of kernels was no longer subjected to his careful crossing with select specimens, they reverted back to normal. Mutations, it seems, would have done the trick where selection had failed. Thus, natural selection was vindicated over pangenesis and neo-Darwinians were quick to recast Darwinian theory by changing variation to mutation. In effect, notes Himmelfarb, natural selection becomes a court of last appeal in the process. When it was discovered that favorable mutations are extremely rare in nature, neo-Darwinists then insisted that this simply goes to prove the very power of natural selection, which is able to surmount such formidable odds. The neo-Darwinians, it is apparent, comments a skeptical Himmelfarb, are as adroit as Darwin in making a virtue of necessity and in converting difficulties into assets (p. 329).

But Darwin also increasingly relied on sexual selection as an adjunct to natural selection, and a considerable portion hisDescentwas spent laying out its features. Here Darwin argued that some traits evolvednotthrough interspecies competition but through intraspecies competition, the selection of mates and breeding those traits deemed most desirable into the species. So while the natural superior strength of the male was obviously derived through natural selection, he attributed the beard to be an ornament to charm or excite the opposite sex. For Darwin music and the sweeter voice of the female were all explained as accoutrements for sexual attraction. Alfred Russel Wallace thought all this sexual selection talk was nonsense. Wallace, who had spent nearly twelve years with indigenous peoples, from South American natives of the Uaups River Valley to Dyak headhunters in Borneo, pointed out to Darwin that tribal women rarely if ever sing and that what an Englishman might value as a sweeter voice was thoroughly uninteresting to the aboriginal peoples he knew. Even if certain cultures could be found where some women did sing, the result would seem to be a zero sum gain for the sexual selection theory and thus no explanation at all. Wallace further pointed out that Darwins suggestion that elaborate plumage in birds gave evidence of sexual selection was, in fact, merely signs of sexual maturity and vigour and that to ascribe the elaborate coloration in butterflies, which is strikingly similar to birds, to female choice unthinkable (see hisThe World of Life). In short, Himmelfarb points out, Sexual selection has all the faults of natural selection and more:the suspicious facility with which it can be made to explain anything and everything, the manipulation of evidence for whatever purposes are convenient, and the invocation of ignorance when all else fails. Ignorance is resorted to even in so crucial a matter as the intellectual disparity between man and the apes (p. 366).

Here again we are treated to a litany of guesses and conjectures. If man is closely allied to the higher Simi, as Darwin openly suggested to the Marquis de Saporta, and is, in fact, descended from some primate ancestor then a naturalistic explanation for the development of speech, that uniquely human characteristic, might go a long way in revealing the process of divergence from our alleged animal ancestors. Darwin suggested that speech and language may indeed account for humanitys great intellectual advance. Here, Darwin suggested, this most distinctive of human attributes probably originated in the imitation and modification of various natural sounds, the voices of other animals, and mans own instinctive cries, aided by signs and gestures. Darwin then leans on sexual selection to invest early man with true musical cadences, that is in singing speculating by a widely spread analogy, that this power would have been especially exerted during courtship of the sexes . . . .10Of course, Wallaces experience among native peoples didnt bear this out, but Darwin used it and maintained it anyway.

But this forced Darwin into an even more difficult conundrum. How could grunts and groans develop into intelligible speech unless a brain sufficiently advanced to develop it already existed? Darwin merely relied upon some early progenitors of man, but wasnt this precisely what Darwin had called upon speech to explain?As for other behaviors deemed utterly counter to the good of the group his shock at the utter licentiousness and unnatural crimes of many tribal cultures Darwin simply explained them away as evidence of their insufficient powers of reasoning. Given all the foregoing, one is forced to agree with Himmelfarb: When there are more exceptions to the rule than exemplifications of it, it would seem time to abandon the rule (p. 373).

Why did Darwin retreat from natural selection into subsidiary notions of pangenesis and sexual selection? Was Darwin simply adrift in theories? Himmelfarb keenly explains:

It was not, however, without cause that he [Darwin] abandoned natural selection. What forced his hand was the realization that natural selection was untenable as the main explanation either for the development of man from the animals or for distinctions of race and sex. Natural selection assumed that beneficial variations alone would be preserved. The difficulty was that the races of man differ from each other and from their nearest allies amongst the lower animals, in certain characters which are of no service to them in their ordinary habits of life. The advantage of sexual selection was that it did not have to prove utility. . . . More and more, the Lamarckian principle of the inherited effects of use and disuse came to replace natural selection (pp. 366-367).

This too would define Wallaces break with Darwin. For Wallace, sexual selection and pangenesis were unnecessary. Instead, he (in true original Darwinian fashion!)stuck with the original formulation: natural selection was guided and directed by the naturalistic principle of utility. But the very thing which defined italsolimited it for Wallace. How could one satisfactorily account for the mind of man? For that matter, what explains the origin of life before there was a principle of utility in operation? For Wallace, the answer was to be found in a teleological universe directed by an Overruling Intelligence. For Darwin, of course, this was unacceptable. But, as Himmelfarbs cogent analysis demonstrated, neither were Darwins wholly naturalistic ones.

Howdowe explain the origin of man? There are really just three options: invoke some form of teleology (by far the general consensus worldwide past and present has been some form of theistic teleology); leave the question unanswered (and some, like philosopher Thomas Nagle, have done precisely that); or rely upon some wholly naturalistic explanation, most commonly Darwinian evolution (a position held by an elite minority only over the past few hundred years). While the first option has usually had an ameliorating effect on the harsh consequences of competition and rivalry and infused society with needed altruism and the second merely provides external critique of proffered explanations, the third has had a dubious history, especially when a theory becomes itself an ideology in the hands of passionate ideologues.

Himmelfarbs chapter on Darwinism opens by observing that when applied to a variety of social contexts it could have a free and loose translation which provided the added advantage of giving it license to a variety of social gospels (p. 412). Applied to many social issues, Darwinism was ambiguous. Darwinism, for example, could argue against slavery, the greatest endorsement of which came from Darwin himself who was an outspoken critic of this peculiar institution. Recently Adrian Desmond and James Moore elevated this to a motivating factor for Darwins theory in theirDarwins Sacred Cause(see review). The thesis is plausible, after all, DarwinsOriginwas written and published when the slavery controversy (which the British Empire had abolished earlier in 1833) raged in America.But as Himmelfarb points out the implications of Darwins evolutionary theory could be taken in other ways:

It was not necessary . . . to confute theOriginin order to justify the South. It was only necessary to re-interpret it. For there were features of Darwins theory that could easily give comfort to the proponents of slavery and racism. Although Darwin derived all races, like all species, from a single historic ancestor, he by no means denied the reality of separate races and species in the present. . . . Nor did he deny that under certain conditions it was desirable to maintain, as far as possible, the purity of races. TheOrigin did declare that crosses between varieties tended to increase the number, size, and vigor of the offspring. But this was true only in special cases: where, for example, the crossed varieties had previously been exposed to fluctuating conditions and thus were especially hardy. Otherwise, such a cross might prove fatal to both varieties.

It was this argument against the crossing of races that first impressed itself upon some of the readers of theOrigin. One month after its publication, on the occasion of John Browns raid at Harpers Ferry, theTimesgave warning that the abolitionists would turn the population of the South into a mixed race. The lesson of modern times, it said, was that such a mixture of races tends not to the elevation of the black, but to the degradation of the white man. Reading this, a secretary at the American legation in London observed: This is bold doctrine for an English journal and is one of the results of reflection on mixed races, aided by light from Mr. Darwins book, and his theory of Natural Selection.

The subtitle of theOriginalso made a convenient motto for racists: The Preservation of Favoured Races in the Struggle for Life. Darwin, of course, took races to mean varieties or species; but it was no violation of his meaning to extend it to human races, these being as much subject to the struggle for existence and survival of the fittest as plant and animal varieties. Darwin himself, in spite of his aversion to slavery, was not averse to the idea that some races were more fit than others and that this fitness was demonstrated in human history (pp. 415-416).

Indeed even Desmond and Moore admit as much. When Darwins friend Charles Kingsley, whose family had been financially ruined when West Indies slaves were emancipated under British law, suggested that the lowly races were doomed and that the white race was destined to domination he was expressing common belief in Victorian England. Even Darwin, confess Desmond and Moore, agreed to the gruesome prospect: It is very true what you say about the higher races of men, when high enough, will have spread & exterminated whole nations. There was a fatalism to the statement. While slavery demanded ones active participation, they add, racial genocide was now normalized by natural selection and rationalized asnatures way of producing superior races. Darwin ended up calibrating human rank no differently from the rest of his society (Darwins Sacred Cause, p. 318). So much for Darwins sacred cause!

It was rationalized in another context as well: From [Darwins] . . . preservation of favoured races in the struggle for life, it was a short step, Himmelfarb points out, to the preservation of favored individuals, classes, or nations from their preservation to their glorification (p. 416). And converselyevenfrom the most favoreds preservation to theleastfavoredselimination. Recent [Himmelfarb is writing in 1959] expressions of this philosophy, such asMein Kampf,are, unhappily, too familiar to require exposition here. And it is by an obvious process of analogy and deduction that they are said to derive from Darwinism (p. 417). (More recently historian Richard Weikart has thoroughly explicated the connection inFrom Darwin to HitlerandHitlers Ethic).

While the Nazi program to eliminate the unfit is an indelible stain on human history, it should be remembered that it was but a specific (albeit especially horrific) exercise of eugenics being applied less dramatically elsewhere, and nowhere more enthusiastically than in the U.S. Described by its leading American apostle Charles Davenport as the science of the improve-ment of the human race by better breeding, eugenics could be applied (as this definition implies) positively by encouraging better marriages and family unions but also negatively as in the culling of the unfit. It was on this basis that more than 65,000 Americans were forcibly sterilized in the early decades of the 20th century. With Indiana passing the first sterilization law in 1907, Californias law of 1909 surpassed all others in efficiency, sterilizing more than 2,500 in ten years and in the next ten years accelerating the effort against the unfit to sterilize an additional 3,500 more. Following World War I, more than twenty states passed sterilization legislation, many modeled on the Indiana and California examples. (For details see Harry Bruinius,Better for All the World: The Secret History of Forced Sterilization and Americas Quest for Racial Purity, 2006. For a more complete and explicit connection between Darwinism and the American eugenics movement see John G. WestsDarwin Day in America.)

The champions of American eugenics were Charles Davenport and Harry Laughlin. Davenport devoted himself to natural science and became committed to the ideas of Francis Galton. It was Davenport who convinced the wealthy widow Mrs. E. H. Harriman to fund the Eugenics Record Office (ERO) at Cold Spring Harbor, New York. In October of that year a Missouri schoolteacher, Harry Laughlin, accepted Davenports invitation to become superintendant of the ERO. In a program described by Bruinius as stunningly ambitious, Laughlin compiled information on existing sterilization laws with the hopes of coordinating a more centralized national legislative plan. A champion of social Darwinism, Laughlin insisted, To purify the breeding stock of the race at all costs is the slogan of eugenics. . . . It is at once evident that, unless this complementary agency, compulsory sterilization of certain degenerates, is made nation-wide in its application, and is consistently followed by most states, it cannot greatly reduce, with the ultimate end of practically cutting off the great mass of defectives now endangering the conservation of our best human stock . . . (quoted inBetter for All the World, p. 212).

In either case whether in Germany or America the connection with Darwinism is clear. Darwin insisted in hisDescent of Manthat humankind differed from animals in degree not in kind. If species are directed by the blind forces of natural selection, then why not give nature a helping hand by moving it along a better and perhaps more direct path to improvement? Many eugenicists reasoned that Darwins own examples of pigeon breeders merely proved the point. Such ideas were promulgated by Darwins cousin Francis Galton. He coined the very term eugenics, meaningeu(good or well in Greek)genes(born or birth). Nevertheless, as indicated above, it was not England that would adopt these ideas programatically. With America leading the way under the missionary zeal of Charles Davenport and Harry Laughlin as mentioned earlier, Hitler need go no further than Laughlins Model Law as an example in framing his Law for the Prevention of Genetically Diseased Offspring in which more than 150,000 Germans were sterilized as unfit. No wonder then that the University of Heidelberg awarded Laughlin an honorary doctorate for his pioneer work in racial hygiene in 1936.

How different was the view of natural selections co-discoverer, Alfred Russel Wallace! Wallace was adamantly opposed to eugenics. In the last book that he would write,Social Environment and Moral Progress(1913),Wallace referred to eugenic proposals for the segregation of the feeble-minded, sterilization of the unfit, and destruction of deformed infants, suggestions in every way dangerous and detestable, and efforts to interfere with the freedom of marriage . . . not only totally unnecessary, but . . . a much greater source of danger to morals and to the well-being of humanity than the mere temporary evils it seeks to cure (pp. 142-143).Wallace further explained (as indeed he had in hisWorld of Life, [1910]) that natural selection, which determined with law-like severity the brutal struggle of species, was no longer applicable to man:

From the moment when the first skin was used as a covering, when the first rude spear was formed to assist him in the chase, when fire was first used to cook his food, when the first seed was sown or shoot planted, a grand revolution was effected in Nature a revolution which in all previous ages of the earths history had had no parallel. A being had arisen who was no longer subject to bodily change with changes of the physical universe a being who was in some degree superior to Nature, inasmuch as he knew how to control and regulate her action, and could keep himself in harmony with her. Not through any change in his body, but by means of his vast superiority of mind (Social Environment and Moral Progress, p. 110).

How this happened Wallace termed the Divine influx, a point in time when by purposeful action some portion of the spirit of the Deity, man became a living soul (p. 102). By limiting natural selection to the principle of utility first enunciated by Darwin himself, Wallace was able to discern discrete examples of intentional design in nature (the most stunning being the human mind) to counter Darwins naturalistically bound biological processes to incorporate genuine theism in a teleologically liberatedintelligent evolution(for details seeAlfred Russel Wallaces Theory of Intelligent Evolution).

This idea was completely rejected by Darwin and his followers. Instead, as seen earlier, Darwin and his disciples speculated on theories of pangenesis and sexual selection. Darwinians will plead that Mendel rescued their evolutionary theory, a curious position given the fact that, whatever the respective merits of the two mens ideas, Mendel himself opposed it and specifically arguedagainstDarwinsOrigin.11So what are we to make of Darwins contribution to the broad history of ideas and to society at large? Himmelfarb concludes her study with a review of the Darwinianrevolution. Some, like Ernst Mayr (see his Darwins Impact on Modern Thought) and more recently Peter Bowler (see his Darwins Originality) believe Darwin to have effected a thorough and sweeping revolution of historic proportions. If one is to measure it by the effect it has had on ethics, morality, and the general secularizing of society its revolutionary impacts seem undeniable. But revolutions dont necessarily imply progress and advance. Here Himmelfarb points out that Darwin was the leader of a distinctlyconservativerevolution. Oswald Spengler thought theOriginreeked of the atmosphere of the British factory (p. 418). The modernity to which Darwin brought the world was built upon foundations long preceding him. Surely the ideas of Ren Descartes and Thomas Hobbes had cleared the ground for the scientistic edifice that Darwin erected after all, Darwinian theory is if nothing else the gospel ofbellum omnium contra omnes the war of all against all that epitomized Hobbes characterization of human existence. And we know that Darwin was familiar with the skepticism of David Hume and the positivism of Auguste Comte. In this sense Himmelfarb is quite correct: Darwin, dramatizing and bringing to a climax the ideas, sentiments, and conjectures of his age, may be thought of as a hero of a conservative revolution (p. 447).

In the end, Gertrude Himmelfarb presents a complete and honest portrayal of Darwin and his theory, and her points are compelling:

If the Darwinian faithful cannot abide the less than ideal portrait that emerges they have only their Down House hero to blame.

Since writingDarwin and the Darwinian Revolution, Gertrude Himmelfarb has moved on to treat a wide range of topics. Nevertheless, her influence as an especially cogent historian of the man and his theory continues. A few have taken notice. Margaret A. Faye, for example, mentions her insightful and lucid analysis.12Philosopher/theologian Edward T. Oakes, S.J., PhD, wrote: I awoke from my own Darwinian dogmatic slumbers only late in life, when I first read Gertrude Himmelfarbstour de forceof a biography . . . .13M. D. Aeschlimans Angels, apes, and men praised her devastating critique for exposing the internal inconsistencies and willful obfuscations that have characterized Darwinism from the beginning, yet noted the conspicuous neglect of her work by those suspiciously interested in promoting the Darwin brand.

Neglected perhaps but not without opportunities for exposition. Four years ago the publication of edited compilations of Darwins works, E. O. WilsonsFrom So Simple a Beginning: The Four Great Books of Charles Darwinand James D. WatsonsThe Indelible Stamp: The Evolution of an Idea, offered treatments by two of this tormented evolutionists most adoring fans and the occasion for a reply by Ms. Himmelfarb.

It appeared as an essay review titled Monkeys and Morals inThe New Republicon December 12, 2005. Pointing out that natural selection rather than evolution was Darwins claim to fame, Himmelfarb states that, interestingly enough, it was secularists who voiced concerns about the theory as much if not more than the religious community. Citing no less than John Stuart Mill, she notes his admission that the theory was impressive enough but that even as late as 1870 he confessed it to be problematical. Instead, Mill concluded that the evidence suggested creation by intelligence:

Creation by intelligence this by Mill, hardly a religious dogmatist. Today one may hear echoes of those words in the theory of intelligent design, which is derided by most scientists (including the editors [Wilson and Watson] of the present volumes) as a euphemism for creation and thus a denial of evolution. And so it is, for some of its proponents. Yet others, themselves scientists, insist that their quarrel is not with evolution itself but rather with natural selection conceived as a purely mechanistic and entirely sufficient explanation for evolution. For them, intelligent design is nothing more or less than teleology, the recognition of a purposiveness or direction in nature, with or without a Creator in the orthodox sense of God (p. 35).

Julian Huxley, Thomas Henrys grandson, thought the modern evolutionary synthesis solved all of Darwins problems. But, as Himmelfarb observes, Notwithstanding Julian Huxley, nothing has been settled. And notwithstanding the editors of these volumes, too, who sometimes sound as dogmatic as the creationists they deride not only in respect to evolution (the blind force, as Wilson puts it, that created animals and man) but in respect to all human behavior (p. 36). The intransigent scientism of both Wilson and Watson is duly noted too, and the lessons of their enormous achievements are countered in perhaps greater measure only by their hubris.

Himmelfarb ends by bemoaning the polarizing effects of both sides of this controversy, though as expected, this seasoned historian casts a far more wary eye on those (like Wilson and Watson) who hold the present positions of power in the debate. If she registers distress over the renewed warfare, a war incorrectly drawn between science and religion by John William Draper and Andrew Dickson White in the Victorian era with attempted truces more recently proposed by Stephen Jay Gould and Ronald Numbers, it would do well for everyone to realize that the battle isnotbetween science and religion butwithinscience itself.

Is methodological naturalism theonlyappropriate avenue of scientific inquiry?Arewe to believe in a uniformity of natural causesin a completely closed system?Ishuman intellectual endeavor easily, and more importantly,properlydivided into Non-Overlapping Magisteria?Is the human intellect reducible itself to purely naturalistic explanations so that our neurons and synapsearewho weare? The problem with the truces of Gould and Numbers ( and indeed with a wide variety of so-called well-intentioned theistic evolutionists like Ken Miller, Karl Giberson, and others) is that they would bury the hatchet only by burying it into the heads of those proposing intelligent design. After all, answering all these questions in the affirmative without substantively engaging in meaningful dialogue winds up merely ratifying the reigning Darwinian paradigm. But the very nature of the questions themselves bespeak the limitation of giving positivistic answers like those attempted by Wilson, Watson, Gould, or Numbers. While Himmelfarb yearns for peace, her suggestion that there need not be any inherent opposition betweentheism and evolution is a sound one. But itcannotbe an evolution hidebound to a conception of science thata prioriprecludes it.

Nevertheless, in providing a bold and brave historical analysis to the question of Darwinian evolution, Himmelfarb has rendered invaluable service. Her incredulity over those who continue to insist that Darwinism is theonlyreceived truth remains a mark of her constancy on behalf of reason and free and open inquiry. Her willingness to swim against the tide of nodding acquiescenceis the measure of a scholar genuinely committed to following the evidence wherever it may lead.

I am happy to report that Professor Himmelfarb remains active and intellectually vibrant and appeared recently onBook TVdiscussing her work on Edmund Burke. While her historical acumen has moved on to treat other subjects, she has indeed made an enduring contribution to our understanding of the figure of Charles Darwin and the modern cultural paradigm of Darwinism. If she suggests that the man should become less iconic or that the theory should be more modest, it is only the counsel of a historian reminding us, as Herbert Butterfield did long ago, that in matters of history as well as in science we should all be a bit less Whiggish in their pursuit.

Photo credit: National Science and Media Museum, via Flickr.

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Himmelfarb and Her Haters - Discovery Institute

These Vaginas Evolved to Fight the Penis, Not Accommodate It – VICE

Evolutionary biologist Patty Brennan had watched a lot of birds have sex. But in 2002, in Costa Rica, she saw something she never had before: a bird penis.

Most male birds don't have penises. They mate using an opening called a cloacaderived from the Latin word for sewer. It's a cavity inside a bird's anus that's a one-stop shop for the digestive, urinary, and reproductive tracts. When birds mate, the male and female cloaca touch. The male releases sperm, and it enters the females body. It's referred to, somewhat romantically, as a "cloacal kiss."

Brennan was observing a pair of Great tinamousbrown, chicken-like birds with small heads that live in the Costa Rican forest. Instead of just the subtle and brief cloacal kiss, the male bird grabbed the female by the neck. Then, the two birds started walking around still attached, as if they were fused together. When they separated, she saw a white, tentacle-looking organ hanging from his body.

"This was unlike anything I had ever seen," she said. "I was like, is this a penis?" (According to biologist Richard Prum, the tinamous penis had been seen and described by Victorian anatomists, but the appendage was forgotten to science. Her sighting was probably the first-ever observation of the tinamou penis in action," he wrote in a 2017 book.)

That unexpected bird penis launched Brennan, now an assistant professor of biological sciences at Mount Holyoke College, into a career of studying the weird and wonderful variations of genitalia in the natural world. But unlike many scientists before her who had noticed the dizzying variety of penises out there, Brennan began to ask: What about the vaginas? A long-standing misconception in evolutionary biology was that penises were incredibly diverse, but vaginas were not. In the past two decades, biologists, like Brennan, have been finding otherwise.

While doing so, they've been uncovering how gender biases might have played a role in obscuring vaginal variety, and how excluding vaginas from the study of genital evolution led to gaping holes in our understanding of why genitals look and behave the way they do. Only by examining how male and female parts evolve together can we see how sometimes, strange genitals are a result of sexual conflicteach sex trying to get the upper hand to control the reproductive act to best suit their needs. That's what Brennan learned, not through bird penises alone, but also through the vaginas with which they interact.

After her encounter in Costa Rica, Brennan wanted to continue studying bird genitalia. She shifted her focus to ducks, a more accessible subject than tinamous. At a duck farm in California's Central Valley in 2009, she captured some duck penises in action. (These ducks had been trained to ejaculate into small glass bottles for artificial insemination.)

You might remember what she and Prum discovered because, for a short while, the duck penis went viral online. Brennan found that the penises unfurl out of a duck's body at lengths of around 5 to 7 inchessome duck penises can be almost as long as the male's body. And they were spiraled, like a fleshy cavatappi pasta noodle. Male ducks forced these long corkscrew penises onto females. The internet was horrified, and also, enthralled.

In the history of people (and scientists) marveling at genitalia in nature, this is where it often stops: Look at this weird penis! In 1979, Science published a paper on the penis of the damselfly. As Dutch evolutionary biologist, Menno Schilthuizen, wrote in his book, Nature's Nether Regions, this minuscule penis carried a miniature spoon that, during mating, cleaned out the females vagina, scooping out any remaining sperm from previous males. It was an eye-opener as well as a sperm-scooper.

This finding opened biologists' eyes to the fact that even tiny creatures had strange penises. The chicken flea's penis is rolled up in its body like a coiled spring. Other insects have musical penises, where males rub them against ribbed parts of their bodies to emit loud noises. Black widow spiders have penis tips that break off to block other males sperm from entering a female. It was somewhat of an evolutionary mystery: Why were penises so different from one another if they had the same evolutionary purposeto deliver sperm to a female's eggs?

The lock and key theory was one potential explanation, proposed in the mid 19th century. It said that male genitals were like a key, and for each key there needed to be a corresponding lock (the vagina). If the key doesnt fit into the lock, mating couldn't take place. Essentially, penises varied to keep different species from mating with one another. Another guess was sexual selectionthat females detected some particular feature of the male genitalia and used it to choose a mate, pushing the male's penis evolution in bizarre off-shoots.

Still, the focus remained on male genitalia and how it was changing and evolving, even in more recent texts on genital evolution, like important work from scientist William Eberhard on sexual selection. Brennan wrote in a 2016 paper that while Eberhard noted female choice was important in shaping male genital features, he concluded that female genitalia are relatively uniform while male genitalia are diverse.

"It created this idea, from my reading of the literature later on, that the females were somehow boring," Brennan said. "We need to look at the males, because thats where all the action is.

As a result, most of the research on genital evolution has focused on males. Nearly two times as many studies have looked at male genitals compared to females. In 2014, evolutionary biologist and gender researcher Malin Ah-King and her colleagues looked at 364 studies published over the last two decades, and found that 49 percent of them only looked at male genitals, compared to 8 percent that looked only at females, and 44 percent that looked at both.

Even the language that researchers use to describe male and female genitals has differed. A study found that active words like "coercion" are used for males, while more passive words like "avoidance" or "resistance," are provided for females. As Ah-King and her co-authors wrote: Too often, the female is assumed to be an invariant container within which all this presumed scooping, hooking, and plunging occurs.

When Brennan first saw the duck penis, though, she immediately considered the duck vagina. I looked at their penis and next question was, 'wow these penises are so big. So what do the vaginas look like?' Surprisingly, no one had investigated that before, she told me. To her, it was an obvious question. As she told science writer Carl Zimmer for a New York Times article: You cant have something like that without some place to put it in. You need a garage to park the car.

When she dissected some female ducks, "I could not believe it, she said. The differences in the vagina of a duck compared to the vagina of a chicken or a finch or quail was like the difference between night and day.

What Brennan found was a vagina like a labyrinth. Yes, duck penises were spiraledbut duck vaginas were too, in the opposite direction. Rather than finding a vagina that had evolved to fit this weird penisa garage that fit the carthe duck vagina indicated a less cooperative history.

Given that duck mating was often forced, Brennan and her colleagues hypothesized that the vagina had co-evolved to actively resist the males. The ducks vagina is swirled in a clockwise coil, so the males can only completely penetrate her with their counter-clockwise penis if she chooses to relax her vaginal muscles. Even though female ducks can't stop the male ducks from forcing themselves on them, they can control if the male could successfully inseminatereclaiming some reproductive autonomy.

Brennan and her colleagues looked at other species where the males took part in forced copulation, and then at the corresponding females. In ducks and geese, they found that when male birds forced sex on the females, females also had complicated vaginas. In species where theres no forced copulation, then the females have a regular, tube-looking vagina," Brennan said.

It also meant that the duck penis size and shape wasn't solely a result of males competing with other males, or females making a choice between malesit was the female and male ducks' competition driving the evolution.

This is the core tenet of sexual conflict: Males and females dont always agree about the best way to mate. For males, mating with a large number of females is the ideal way for them to procreate. For females, who are often left with the care of the offspring, as well as giving birth and pregnancy, being selective about reproduction is her best bet for creating progeny that will survive. This creates a conflict, where the males are going for quantity and the females, for quality.

Lets say a male animal evolves a penis hook, which allows him to latch onto a female. Even if that hook hurts the female, or gives her an infection, if it benefits the male by allowing him to reproduce more, the genes for that hook will be passed to the next generation. That puts the female a step behind, so evolution might next select for females that can defend themselves against the hook, and evolve thicker walls in their vagina. (Something very similar has happened in sharks.) This is a way of understanding the evolution of genitals as a kind of conversation, even if a contentious and competitive one. And this perspective is providing new understanding for a whole host of creatures.

The males evolve these weird penises and females evolve their convoluted vaginas in response, Brennan said. This is a lot more widespread than what we had originally realized. It's just, we have to go out there and look.

Take the earwig, an insect with a male reproductive organ called a virga. The virga has a fringe-like tip that can brush away sperm from any male that mated with a female before him. Looking at the male genitals only tells one half of the story, because the females have receptacles in their bodies to store sperm that lie just out of reach of the virga. It may seem that the males are controlling the sperm, but the females have the upper hand. As science writer Ed Yong wrote: The male can scrape away all he wants; the female decides whether to keep or jettison her sperm.

Dolphins have a complex series of vaginal folds that researchers once assumed were there to keep sea water from getting inside the female reproductive tract. Theyre realizing now how intricate their vaginas are, partly by making the effort to look closer at them. In 2017, biologist Dara Orbach made silicone molds of the dolphin vagina "revealing complex folds and spirals," the New Scientist reported. Brennan said it's now thought that those folds are actually barriers to male's penises.

Paying more attention to vaginas can help explain strange mating behavior too: In water striders, bugs that live and walk on water, the females evolved a genital shield, which can block any males that try to force them into mating. That led the males to adopt new "courting" techniques. "The males have started tapping the surface of the water while mounted on a female; the resulting ripples attract fish, and since the female is under the male, she's more likely than him to become a meal," according to post on Nature's blog. "Females can avoid this grisly end by giving in to the male's intimidation and mating with him.

Without knowing that the females have a genital shield, researchers' understanding of such behavior would be incomplete. It allows us to understand all of these bizarre morphologies and behaviors that we see in the context of, essentially, an arms race, said Teri Orr, a evolutionary ecologist at The University of Utah.

Spiders are another of Orrs favorites, because they can have around a dozen different pockets in them for manipulating spermsome are for receiving sperm, or moving it around. Orr frequently studies bats, and said they will store sperm for a full year in the reproductive tract. Leaf cutter ants can store sperm for around ten years.

Female chickens can eject about 80 percent of sperm from undesirable mates. Female guppies can hold onto sperm tooone study found that one in four guppies in Trinidad and Tobago were fathered by males that had been dead for 10 months. By doing so, females could wait to reproduce at favorable times of the year.

Theyre able to keep those sperm until its a good time of year for them to become pregnant, and then carry out that pregnancy and have babies when theres food available for them, Orr said. To me, that is absolutely mind-blowing. A lot of it is almost science fiction, what these species are able to do."

It also shows how the female anatomy is anything but passive. Outside of sperm storage, the vagina is awash with muscles that control contractions and movementits as mobile as the digestive tract is, Orr said. These muscles can play a part in moving the sperm where they want it to go. We didnt know what until about a decade ago, she said. And even then, its only in cattle, horses, mice and humans that its been studied. Thats such a small part of the diversity thats out there.

In 2005, more than 200 scientists met in London at The Royal Society for a meeting titled Sexual conflict: a new paradigm? Brennan said that since then, she feels the field is moving to include vaginas, and that several of the most recent papers on genital evolution acknowledge the fact that female genitals have been overlooked. But Orr said that when she presents her work at conferences, it can still feel like its regarded as out there" or niche. It hasnt reached mainstream science yet, she said. I think its going to take a little while until its fully embraced and not just a noveltybut normal biology.

It's not as if Brennan wants the research to flip and only focus on femalesthe point is that you need both pieces of the puzzle. Ive been very adamant that when youre looking at genitalia, you cant just look at the female or the male alone," she said. "You need to look at both because of that mechanical fit. I could commit the opposite sin, in a way. I could just go look at a bunch of females and never look at the males. Thats not going to tell me much.

She hopes that the field of genital evolution become more well-rounded, and also that the public will recognize its value. When Brennan's work on duck genitalia went public, conservatives latched onto it as a waste of government money (like a lot of academic research, it was partly funded by the National Science Foundation), acquiring the moniker #DuckPenisGate. Fox News put up a poll on their site where readers could vote if the research was a worthwhile use of taxpayer money, and 89 percent voted it was not. Brennan and her co-author Prum had to write articles defending the research.

The thing about basic science, Brennan said, is that you never know when a seemingly obscure discovery in nature is going to lead to an application for humans. So could secrets to our evolutionary past (and sexual conflicts) be hidden in our genital shapes? It's true that humans also have weird penises awash with unsolved questions, Brennan said. They are disproportionately wide given our body size and MRI studies of people having sex show that the shape of the male and female genitalia can change during intercourse, for reasons that are not completely understood.

Humans don't have penile spines, unlike many of our close primate relatives. Humans have also lost the baculum, a little bone inside of the penis of many animals, like bats, rodents, and primates. We have no idea what it does, Orr said. Its buried in tons of soft tissue and so its not interacting with the female, so its quite mysterious." Even less understood is the tiny little bone some animals have inside of the clitoris which humans didn't retain.

But more often, translation from basic science will come from where you least expect it. One obvious example is how the immune system of a bacteria was developed into a revolutionary gene editing techniqueCRISPR/Cas9.

In the realm of genital evolution: duck penises grow and shrink every season, which means there are probably stem cells in the penis that allow for that growth each year. If researchers could learn what those cells are and how they work, they could have all sorts of medical or cosmetic applications. Could we actually grow penile cells that might become a treatment someday? Its perfectly possible," Brennan said.

Many of the stages where pregnancy fails in humans are the same ones where bats are able to intervene and store fertilized eggs or sperm. By looking closer at those processes, it might lead to ideas for aiding issues in human or reproduction or endocrinology, Orr said.

Hypospadias is a birth defect leading to a malformed urethra; one in every 200 boys is born with some type of hypospadias. For people with such developmental problems, or others, like malformation of the uterus, research into genitals that are naturally bifurcated could lead to an understanding of what causes those hiccups, and how to fix them.

Even if those animal-human translations aren't right around the corner, the field of genital evolution has already offered something else: Recognizing the value in seeing how gender biases and language can divert research to ignore crucial elements. Anthropologist Emily Martin's 1991 essay The Egg and the Sperm highlighted how the (often incorrect) descriptions of human sperm and eggs reflected stereotypical male and female roles. It's a reminder that it could happen again, and to examine what social constructs are currently inseminating scientific research.

And Brennan wonders if the response to her research doesn't betray how touchy and judgemental people still are about genitalia, especially vaginas. It's almost as if there was something a little perverse with that line of questioning or that particular type of research," she said. "I happen to think that we actually need to understand a lot more about sex and sexual interactions than less.

She views genitals just like any other organs. If you think about our other organs: livers, kidneys, hearts, or brainsthere's much less variation and excitement. It's a rare window into what evolution can do. Genitalia are critical biological organs to be studying, she said. Im still surprised that we know as little as we seem to know. Evolutionarily, this is where the rubber meets the road.

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These Vaginas Evolved to Fight the Penis, Not Accommodate It - VICE

Whales Help Explain the Evolutionary Mystery of Menopause – WIRED

There's a rare human trait that doesn't often make it into debates about what makes our species unique: menopause. Humans are among just a handful of species where females stop reproducing decades before the end of their lifespan. In evolutionary terms, menopause is intriguing: how could it be advantageous for reproductive ability to end before an individual's life is over?

One possible answer: the power of the grandma's guidance and aid to her grandchildren. A paper in PNAS reports evidence that supports this explanation, showing that killer whale grandmas who have stopped reproducing do a better job of helping their grandchildren to survive than grandmothers who are still having babies of their own.

Its Not All About the Babies

The engine of evolution is offspring. In simple terms, the more babies you have that survive, the more your genes are passed on, and the better the chance of the long-term survival of those genes.

ARS TECHNICA

This story originally appeared on Ars Technica, a trusted source for technology news, tech policy analysis, reviews, and more. Ars is owned by WIRED's parent company, Cond Nast.

But there are other ways to improve the long-term survival of your genes, and that's where evolution gets a little bit more complicated than just brute-force reproduction. If you invest in your siblings' children, or your children's children, you also improve the survival of the genes you share with them. Like every other survival problem that a species must overcomefood, safety, finding a matethe dynamics of natural selection generate different solutions to the question of how to propagate your genes.

The "grandmother hypothesis" suggests that grandmas play a crucial role in the survival of their grandchildren, which obviously gives the grandmas' own genes a boost. But that doesn't explain why humansalong with killer whales, short-finned pilot whales, belugas, and narwhalsstop reproducing with decades left to live. Wouldn't it be better to just keep having babies of your own and help your grandchildren? Possibly not: in certain species, with certain family dynamics, evolutionary models show that it's more worthwhile for grandmas to invest all their resources in their grandchildren, rather than compete with their own daughters.

There's evidence from humans to support this: the grandchildren of post-reproductive grandmas get a survival boost. But there hasn't been any direct evidence of a post-reproductive grandma benefit in other species that have menopauselike killer whales. Similarly to humans, female killer whales stop reproducing around their late 30s or early 40s but can continue to live for decades after that point. Do killer whales also give their grandkids a boost?

Grandma Has Tricks Up Her Sleeve

Like humans, killer whales live in intensely social family groups. Also like humans, young killer whales need help finding food even after they've been weaned. This means an important role for grandmothers, who can share food with their grandchildren and also impart their decades of accumulated experience and wisdom by guiding their families to historically successful feeding spots.

To test whether post-reproductive killer whale grandmas improve the survival of their offspring, a group of researchers collected data on killer whale populations off the coast of Washington state and British Columbia. They tracked the interactions between hundreds of individual whales, recording births and deaths and controlling for the all-important environmental factor of salmon abundance.

Just like humans, whales can become grandmothers while they're still having babies themselves. Because they were interested in the effects of menopause, the researchers wanted to compare the effects of grandma whales that had stopped reproducing to those that were still having their own offspring.

The results showed that grandma whales played a significant role in the survival of their grandchildren. Survival rates dropped sharply for whales that had recently lost a grandmothereven adult whales of 15 or 20 years old. And this effect was more marked when the grandmother was no longer reproducing herself. It was also more extreme when salmon abundance was lower, suggesting that the ecological knowledge of grandmother killer whales is a crucial resource for their families.

Less Competition

This result ties in well with previous evidence on menopause in killer whales, which found that menopause meant a reduction in competition for resources between grandmas and their daughters.

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Whales Help Explain the Evolutionary Mystery of Menopause - WIRED

The Evolution of The Lil Smokies – Flathead Beacon

A few days before his show in Whitefish, dobro player and vocalist Andy Dunnigan of the Montana bluegrass band, The Lil Smokies, traveled across international borders from Vancouver, Canada, a place hes recently started calling home base.

The rest of the band arrived in waves from Oklahoma, Seattle and Missoula to converge together in Whitefish and play a dynamic, triple-header at the Great Northern Bar in Dunnigans hometown.

After performing 160 captivating shows nationwide in the past year, the band was burnt out from traveling. In 10 years of playing bluegrass, The Lil Smokies gradually realized they dont need to live in the same place to be a successful band, and its helped relieve some stress.

In addition to their tour, the five-piece band converged last year to create their new album, Tornillo, named after the small town in Texas where it was recorded.

The recording process differed from the groups previous album, Changing Shades, recorded at SnowGhost Music in Whitefish. For nine days, the band members lived at the studio in Tornillo. They slept there, ate huevos rancheros every morning and immersed themselves in their music in the small town outside of El Paso. The dobro, guitar, fiddle, bass and banjo were their focus.

Because it was so immersive, we had a lot of time to tinker, Dunnigan said.

Before The Lil Smokies arrived in Tornillo, they were fried. In the last few years, theyve grown from a local Montana band into a well-known presence in the bluegrass world. Tornillo brought them back together after years of success, traveling and stress.

It was a very unifying experience, he said.

The recording came at a time of evolution for the The Lil Smokies. Since starting out as a bluegrass garage band playing keggers in their hometown of Missoula, their lyrics have grown deeper and the members more versatile.

Each member has brought a different nuance, Dunnigan said. Were all a little more conscious of space and not as fast and furious as we were. I think thats a sign of an evolved musician. Let the space between the notes prosper.

Tornillo opened the doors they needed to grow musically. The experience allowed them to experiment and impacted them so much they named their album after it.

Before The Lil Smokies were recording albums and playing 160 gigs a year, the band formed during their years at the University of Montana in Missoula. It all started in 2009 when Dunnigan brought along his dobro to a house party, jamming into the night with other like-minded musicians.

We shared a catalog of interest, he said.

From there, they grew as a local Missoula favorite and began playing small gigs in town. It wasnt until they won a band competition at the Northwest String Summit in 2013 and another at Telluride Bluegrass Festival in 2015 that their success skyrocketed. Dunnigan says that was the catalyst that ushered the band onto the road.

Although two members left the group once they started getting big, the newest members, Matthew Rieger or Rev and Jake Simpson, joined three years ago and maintained the Montana vibe.

While they have evolved into a nationwide bluegrass staple, nothing compares to playing in their home state of Montana, Dunnigan said. After the hustle and bustle of the big cities they visit on tour, the band is always excited to come home.

I love the slow pace and the friendliness of Whitefish and Missoula, Dunnigan said. Thats foreign to a lot of places.

Dunnigan says he likes to finish tours in Montana because of the sense of homecoming. Following months on the road, the band is warmed up and ready to showcase their music at home, he said.

The Lil Smokies sometimes make it to play in Billings but always hit Bozeman, Missoula and Whitefish on their tours.

Missoula, Whitefish and Bozeman is the Holy Trinity for us, he said.

A final weekend in Whitefish and a New Years Eve celebration at The Wilma in Missoula is becoming an annual tradition.

Its nice to have these staple events to look forward to every year, Dunnigan said.

The Lil Smokies will finish their tour in Whitefish at the Under The Big Sky Festival on July 18 and 19.

To order The Lil Smokies new album, Tornillo, visit http://www.thelilsmokies.com.

maggie@flatheadbeacon.com

If you enjoy stories like this one, please consider joining the Flathead Beacon Editors Club. For as little as $5 per month, Editors Club members support independent local journalism and earn a pipeline to Beacon journalists. Members also gain access to http://www.beaconeditorsclub.com, where they will find exclusive content like deep dives into our biggest stories and a behind-the-scenes look at our newsroom.Join Now

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Breakthrough in Understanding Evolution Mitochondrial Division Conserved Across Species – SciTechDaily

Cellular origin is well explained by the endosymbiotic theory, which famously states that higher organisms called eukaryotes have evolved from more primitive single-celled organisms called prokaryotes. This theory also explains that mitochondriaenergy-producing factories of the cellare actually derived from prokaryotic bacteria, as part of a process called endosymbiosis. Biologists believe that their common ancestry is why the structure of mitochondria is conserved in eukaryotes, meaning that it is very similar across different speciesfrom the simplest to most complex organisms. Now, it is known that as cells divide, so do mitochondria, but exactly how mitochondrial division takes place remains a mystery. Is it possible that mitochondria across different multicellular organismsowing to their shared ancestrydivide in an identical manner? Considering that mitochondria are involved in some of the most crucial processes in the cell, including the maintenance of cellular metabolism, finding the answer to exactly how they replicate could spur further advancements in cell biology research.

In a new study published in Communications Biology on December 20, 2019, a group of scientists at Tokyo University of Science, led by Prof Sachihiro Matsunaga, wanted to find answers related to the origin of mitochondrial division. For their research, Prof Matsunaga and his team chose to study a type of red algathe simplest form of a eukaryote, containing only one mitochondrion. Specifically, they wanted to observe whether the machinery involved in mitochondrial replication is conserved across different species and, if so, why. Talking about the motivation for this study, Prof Matsunaga says, Mitochondria are important to cellular processes, as they supply energy for vital activities. It is established that cell division is accompanied by mitochondrial division; however, many points regarding its molecular mechanism are unclear.

This exciting new research describes how mitochondrial replication is similar in the simplest to most complex organisms, shedding light on its origin. Credit: Tokyo University of Science

The scientists first focused on an enzyme called Aurora kinase, which is known to activate several proteins involved in cell division by phosphorylating them (a well-known process in which phosphate groups are added to proteins to regulate their functions). By using techniques such as immunoblotting and kinase assays, they showed that the Aurora kinase in red algae phosphorylates a protein called dynamin, which is involved in mitochondrial division. Excited about these findings, Prof Matsunaga and his team wanted to take their research to the next level by identifying the exact sites where Aurora kinase phosphorylates dynamin, and using mass spectrometric experiments, they succeeded in identifying four such sites. Prof Matsunaga says, When we looked for proteins phosphorylated by Aurora kinase, we were surprised to find dynamin, a protein that constricts mitochondria and promotes mitochondrial division.

Having gained a little more insight into how mitochondria divide in red algae, the scientists then wondered if the process could be similar in more evolved eukaryotes, such as humans. Prof Matsunaga and his team then used a human version of Aurora kinase to see if it phosphorylates human dynaminand just as they predicted, it did. This led them to conclude that the process by which mitochondria replicate is very similar in different eukaryotic organisms. Prof Matsunaga elaborates on the findings by saying, Using biochemical in vitro assays, we showed that Aurora kinase phosphorylates dynamin in human cells. In other words, it was found that the mechanism by which Aurora kinase phosphorylates dynamin in the mitochondrion is preserved from primitive algae to humans.

Scientists have long pondered over the idea of mitochondrial division being conserved in eukaryotes. This study is the first to show not only the role of a new enzyme in mitochondrial replication but also that this process is similar in both algae and humans, hinting towards the fact that their common ancestry might have something to do with this. Prof Matsunaga concludes by talking about the potential implications of this study, Since the mitochondrial fission system found in primitive algae may be preserved in all living organisms including humans, the development of this method can make it easier to manipulate cellular activities of various organisms, as and when required.

As it turns out, we have much more in common with other species than we thought, and part of the evidence lies in our mitochondria!

Reference: Cyanidioschyzon merolae aurora kinase phosphorylates evolutionarily conserved sites on its target to regulate mitochondrial division by Shoichi Kato, Erika Okamura, Tomoko M. Matsunaga, Minami Nakayama, Yuki Kawanishi, Takako Ichinose, Atsuko H. Iwane, Takuya Sakamoto, Yuuta Imoto, Mio Ohnuma, Yuko Nomura, Hirofumi Nakagami, Haruko Kuroiwa, Tsuneyoshi Kuroiwa and Sachihiro Matsunaga, 20 December 2019, Communications Biology.DOI: 10.1038/s42003-019-0714-x

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Mary Cariola announces evolution of the organization’s name and identity – WXXI News

The Mary Cariola Childrens Center is announcing a name change as the organization works to evolve its brand.

The non-profit, which provides education and other services for people with disabilities will now be known as Mary Cariola Center, Transforming Lives of People with Disabilities.

President and CEO Karen Zandi says the change reflects the fact that the organization also provides services to adults in residential and community services programs.

Some of that has evolved because of the changes at the state level and individuals who we serve that have multiple and complex disabilities are really pretty challenge and need some individualized plans. So they tend to stay with us until were able to secure other services for them, Zandi explained.

Zandi says Mary Cariola has been serving adults as well as children for some time now.

"Once someone is over 21 talking about them as a child is really disrespectful and therefore deserving the respect that adulthood brings. So the individuals over 21 that we were serving, we didnt want to keep calling them kids," Zandi said.

Mary Cariola, based on Elmwood Avenue in Rochester, serves nearly 500 students from ages 3 to 21 in an 11-county region.

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Mary Cariola announces evolution of the organization's name and identity - WXXI News

The Evolution of Language, by Kinnary Patankar, Henrietta Barnett School – This is Local London

From peng to bants, modern slang has taken on a life of its own. It has even been referred to as its own language, incomprehensible to the adult members of society. English has evolved throughout the country to result in an entirely new set of vocabulary with its own new and distinct definitions. This evolution of language has been happening for millennia, and will certainly continue to occur. From Latin to Old English to slang, humans have refined their methods of communication to involve today over 170,000 words, and across the globe thousands of languages have emerged and developed too. However this begs the question: how did language come to exist in the first place?

It is almost unanimously agreed that it all began with the grunts and noises of cavemen sitting around a fire, not unlike the communication we see today in pack animals such as wolves. The need for language arose from the need for communication; humans have always been social animals and not only crave but require co-operation among a group in order to survive. This line of thinking has resulted in a theory of how language evolved: natural selection. It is known that our brain size is significantly larger than that of our predecessors, and therefore this allowed more complex cognitive function to occur, including language. The humans better able to communicate with each other could find water faster, hunt more successfully, utilise tools together to build shelter. Therefore they outcompeted those unable to do so, and as our brains evolved, as did our language until eventually it comprised of modern day complex constructions such as tense markers, relative clauses and complement clauses. One of the most fascinating aspects of this is that even a young child can learn to speak simply by listening!

This process of language evolution may have begun from simply a grunt to signify that a deer is nearby. However gradually a protolanguage that was made up of rudimentary phrases may have developed, involving stringing multiple words together, such as deer ahead. This phrase contains only two words but is vastly more effective than simple deer as it details its proximity to the hunters. In fact this type of protolanguage can be seen in two-year olds and in the initial efforts of adults attempting to learn a new language. Therefore this theory is very plausible.

On the other hand some argue that language is a result of exaptation: an adaptation being used for something that was not its original purpose, or in very simplified terms, it was a side-effect of other evolutionary advancements in the human brain. These scientists argue that as natural selection favoured adaptations more directly advantageous in the brain such as the abilities of tool making and rule learning, the areas that developed to enhance these functions just so happened to also advance the ability of being able to use language. These two theories are not mutually exclusive, though. They may have occurred in conjunction with each other, allowing humans to begin to use language.

However it is difficult to even agree on what was the beginning of language. Some consider the proto-language, which was used by the Homo habilis, the first language. Conversely others credit it to the Homo erectus. Many even claim that the first language surfaced much later, with the earliest groups of Homo sapiens. Tracking the evolution of language is also incredibly difficult, as it has no physical manifestation, although some fossil records are used to track the shape of the vocal tract (the mouth, tongue and throat). Before around 100,000 years ago, the vocal tracts did not allow the range of sounds and movements of the mouth used in speech today. However this does not mean language started then, as earlier language may have developed without the use of certain vowel sounds and consonants.

Another route scientists are taking is studying other primates, particularly apes methods of communication to perhaps see what had happened to our own species in the very early stages of the development of language. However teaching apes the human language has produced intriguing but only rudimentary results. Ultimately, we are the only organisms with this complex and highly effective method of communication: an ability so unique and demanding such rigorous intellect that it very well may have played a crucial role in humans occupying the role they do today in the natural world. So perhaps the next time you make use of a slang word, perhaps give some thought to how that word came to be in the first place!

By Kinnary Patankar

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The Evolution of Language, by Kinnary Patankar, Henrietta Barnett School - This is Local London

‘Titanfall’ to ‘Star Wars’ to VR realism, the evolution of Respawn games – Los Angeles Times

If anything should put Vince Zampella at ease, one would assume it would be the topic of video game warfare. Zampella, one of the most recognizable figures of the modern gaming era, is after all responsible for much video game carnage.

With his Respawn Entertainment, hes been an architect of sci-fi shooter Titanfall. Previously, with Infinity Ward, he helped define the Call of Duty franchise before an acrimonious split with Activision. Even earlier, with the studio 2015, he contributed to the Medal of Honor franchise, which often features front-line battle action. And yet five minutes into an interview Zampella is struggling to hold back tears as he recalls a visit to Arlington National Cemetery.

What sparked the memory was a seemingly straightforward question about Respawns reboot of Medal of Honor, a release that will essentially take Zampella back to his roots. Had to start with this one, he says, briefly burying his face in his hands.

Its not just that the stakes for the upcoming Medal of Honor: Above and Beyond are higher than when he worked on the 2002 editions of the series the company, after all, is working in virtual reality, where the challenge of palatable video game killing is upped when the participant is in a surprisingly convincing, full-scale virtual expanse.

Rather, Respawn, in reviving the long-dormant Medal of Honor franchise, has several ambitious ideas for adding heft to interactive entertainment. One of them is to intermix very real World War II stories with gameplay, alternating tense, sweat-inducing missions one in which well infiltrate a Nazi palace to steal and burn documents with documentary footage of war veterans retelling and revisiting WWII stories and sites. One such film follows an American veteran in his 90s as he returns to the European battlefield where his friend fell. We watch as he shares hugs and stories with the family that now lives on the grounds, and cameras follow as he leads us to how and where he found his compatriot.

Zampella knows the power of these scenes, having accompanied filmmakers on some of the trips. I took my son with me, who was 18 at the time, and we were at Arlington Cemetery on Gold Star Mothers Day, which is for mothers who lost their children in service. Seeing kids who were my sons age ...

Zampella begins to trail off as his eyes start to water, but he wont be fully derailed. His somewhat guarded personality soon takes over to push back any potential tears and get back to a discussion of what turned out to be Respawns most expansive year. In February, the Chatsworth-based studio surprise-released the instantly popular free-to-play game Apex Legends, which has been sampled by more than 70 million players. Then in November the studio issued Star Wars Jedi: Fallen Order, a return for the space opera franchise to the narrative-driven home video game console, and arguably the least divisive, mass-marketed Star Wars product unleashed this year.

Earlier in 2019 Zampella predicted that Fallen Order, under the direction of Stig Asmussen (God of War III), would begin to remold Respawn developers long typecast as multiplayer shooter guys into interactive storytellers. Now hes ready to look beyond Respawn, the studio he founded in 2010 with Jason West which was acquired in 2017 by Electronic Arts. In 2020 Zampella will also lead the L.A.-based offices of another Electronic Arts-owned studio, DICE.

DICE was founded in Stockholm in the early 1990s and is home to games such as Battlefield, Mirrors Edge and Star Wars: Battlefront. But Electronic Arts characterizes its L.A.-based outpost as long dedicated to support for DICEs core products. Under Zampella, there are plans to expand and launch an original, as-yet-unrevealed game. The company will remain separate from Respawn and, Zampella says, likely will drop the DICE name.

We will probably rebrand, Zampella says. We want to give it a new image. We want people to say, This is a destination you can go and make new content. I think theyve kind of gotten the branding that they are the support studio for DICE Stockholm. I think rebranding is important for showing people, Hey! Come work here. Were going to do some amazing things.

The studio, he stresses, will be separate from DICE Stockholm and separate from Respawn.

Says Zampella, We do talk a lot, and the more we interact and learn from and teach each other, the better well be. So theres interaction, but as far as the games, theyre their own studio.

The move is being overseen by Laura Miele, who about 18 months ago took over as Electronic Arts chief studios officer. At a time when major console makers such as Sony and Microsoft are readying new systems for release in late 2020, and streaming and subscription services are providing more avenues to discover games, Miele says she is focused on broadening the Electronic Arts portfolio.

I think under Vinces leadership the expectation is to have them work on and create a game on their own, Miele says. And I genuinely believe that he is going to help guide them creatively. Hes going to help them further fortify and build out their talent and their team. I think were going to have a really strong studio out of our Los Angeles location. They can go from a support team to a full stand-alone studio to create a new game offering.

Chad Grenier, left, Stig Asmussen and Peter Hirschmann, heads of studio at Respawn Entertainment.

(Mel Melcon / Los Angeles Times)

As for what this means for Respawn, Zampella speaks of his role there now as more of a head coach.

Star Wars Jedi: Fallen Order leader Asmussen will continue to direct a narrative-driven branch at Respawn. The ongoing Apex Legends is overseen by Chad Grenier, who followed Zampella from Infinity Ward to Respawn to work on what would become the first-person-shooter series Titanfall. The virtual reality Medal of Honor: Above and Beyond is being spearheaded by Peter Hirschmann, whose Medal of Honor experience stretches back to the games early days when it was developed by DreamWorks Interactive.

There might be more, says Zampella, narrowing his eyes somewhat cryptically (the company even declines to say how many staffers it currently employs), but he adds, I think my role is the same.

Obviously with Jedi coming out, my focus was a little more on getting that game out the door. Now that its done and out there, its, OK, whats next? Peter will get the Eye of Sauron.

The move to bring in Asmussen was instrumental in changing the tenor of Respawn. Jason West, who recently joined Fortnite developer Epic Games, left Respawn for unspecified family issues in 2013, before the company had issued its first game. Titanfall in 2014 was largely dedicated to fine-tuning the first-person-shooter by creating a game that nimbly moved among different action styles.

Talks with Asmussen began around that same time, before Respawn was acquired by its early investor Electronic Arts, and also before the industry would heavily pivot from single-player adventures to ongoing multiplayer games such as Apex Legends, a Fortnite competitor that lives as a live service with regular updates.

In 2014, I dont think we were talking about single-player action-adventure games being an endangered species, Asmussen says. Thats something that happened while we were working on the game. There was a rise in game experiences that were service-based and people could continue playing over and over. But we stuck to our guns. This is the type of the game we were going to make. I think what happened is live service games expanded the market, but they werent taking away the players who wanted to play a single-player action-adventure.

Star Wars Jedi: Fallen Order is set before Luke Skywalker arrives to destroy the Death Star. Its tale is of a lone, rebellious Jedi. Shown is Jedi Cal Kestis on a zipline.

(Lucasfilm/Respawn/Electronic Arts)

While Electronic Arts hasnt released official sales figures, Fallen Order was tracked as the No. 2 selling game of November, behind Call of Duty and ahead of Pokmon Sword/Shield, according to research firm the NPD Group. Where Asmussens team goes next, he says, isnt set in stone, but there should be plenty of ideas to pull from. Before Respawn knew it was working on a Star Wars game Electronic Arts possesses the license for the Lucasfilm/Disney brand it had developed about 12 prototypes for a new intellectual property, one of which was properly demoed and shopped around the industry.

And Asmussen is steadfast in the types of games he wants to direct. Im not a very good competitive player, he says. Thats when you see the worst of me, in terms of temper. I havent done it in so long because it was getting the better of me. Its a frustrating experience for me. I dont have the patience.

Miele cites Respawns ability to move quickly on an idea to have, for instance, a rudimentary prototype up and running under Asmussen or to shift from a proper Titanfall project to put resources into what would become Apex Legends as a trait she admires in Zampellas teams and wants to spread among Electronic Arts other studios. I feel really strongly about this, she says, and Vince has taught me a lot about this: rapid prototyping.

As I have partnered with him, Ive noticed they get their games stood up and they have hands on their games really soon, sooner than other studios we have, outside of sports, she says. So weve adopted and brought in new prototyping tools and are highly encouraging teams to prototype and prove out game type, game flow and game features before we get to art execution and make it look pretty for executive presentations.

While it may not be apparent from the outside, Zampella says theres a sense of experimentation driving Respawns decisions, even behind more seemingly direct business-driven moves, such as the jump with Apex Legends into the free-to-play battle royale arena popularized by Fortnite.

Zampella and Grenier say theyre not terribly interested in challenging Fortnite as a communal hangout space the latter recently has had crossovers with Marvel and Lucasfilm properties, even hosting a live event with The Rise of Skywalker director J.J. Abrams but they are using Apex Legends as a way to better develop characters, ensuring that game mechanics and art style coalesce into visual storytelling opportunities.

We tend to rapidly prototype characters at a high rate and throw them away when theyre not working, says Grenier. We do 10 times more prototypes than what actually make it into the game.

Or as Zampella puts it, Titanfall, like a lot of other properties in the sci-fi space, started to read as generic sci-fi. He says, Thats terrible for me to say about my own game, but we wanted something more identifiable by going into characters. ... To identify with a character, either by play style or because you like who they are and what they stand for, it was important for us to take that next step. Character-based is now super important to what we do.

Other questions lie ahead. Its clear Zampella has ideas hes not yet sharing on where he intends to take the DICE L.A. team, but Medal of Honor: Above and Beyond will present more near-term challenges. Set in the early 1940s and in development with Facebook-owned Oculus, the game will capture the realistic tone of the early Medal of Honor games each virtual gun, for instance, comes with a very specific set of hand gestures when it comes to reloading.

Shooting in VR is not always a comfortable gaming experience. As fidelity gets better and VR gets more immersive, you kind of feel like youre there. That translates to, Am I harming another more realistic-looking human? Thats something were going to have to be very wary of, Zampella says. When you know the setting is life-and-death and its a historical thing while you may be causing harm to virtual humans youre doing it for the good of other virtual humans in that simulation its something that was valuable to the world.

Such questions fuel Respawns desire to connect in-game conflict to real-world stories and people, says the games lead Hirschmann. We just got a rough cut of a guy named Frank who served in the Pacific, and the stories he tells about what he went through in the submarine service are just crazy. So again, we try to help it hit home that this really happened. These were 19-year-old kids. And you know, often thats our target audience. So its always good if we can build empathy and ignite peoples imaginations. Then maybe theyll come away understanding the conflict a little more.

Its clear Zampella thinks the studio is up to the challenge. The only time he bristles during an interview is when asked about giving up Respawns independent status to sell to Electronic Arts. While the latter provided some of the seed money to help launch Respawn, Zampella also spoke in the early 2010s about the creative importance of maintaining some self-sufficiency from corporate overlords.

Did I give up my independence? he shoots back. Hold on. This is new news.

He continues, For a small studio to do as much as I want to do, it made sense for us to join forces with EA. In talking to EA, they wanted the influence of me coming in to help shape the future of EA. The industry is changing, and we have the chance to be at the forefront of that. Being able to take on new challenges, like DICE L.A. falling under me now, is exciting. I want to challenge myself. I want to do something bigger and funnerer.

And to ensure the conversation ended on a lighthearted note, he added: Use that word: Funnerer.

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'Titanfall' to 'Star Wars' to VR realism, the evolution of Respawn games - Los Angeles Times

The evolution of Sicilian weddings – Pocono Record

Every winter I take a vacation abroad, and I always try to crash a wedding if possible. Preparing in advance I want to learn about the local traditions. This year its Sicily, and as you read this Im there right now!

As I studied up on their wedding traditions Ive learned that for Sicilians, a wedding is probably the most important event of a lifetime. Historically it was a necessity adding to social status, bringing the couple into the community. Unmarried people were considered weird, unlucky and not worthy of attention.

As in other parts of the world, marriages were mostly arranged by parents or relatives, and couples often married without even knowing one another.

Christianity was established in Sicily in the 5th century. The Byzantine, or Orthodox Church became their tradition, but by the 13th century the Roman Catholic Church made inroads on the island.

I found this tidbit to be fascinating: many Sicilian wedding customs, especially before the 20th century, were based on Muslim practices dating back to medieval times when Arabs dominated the island. The church may have supplanted the mosque, but many Muslim traditions held on. It is not uncommon around the world to find subtle influences rooted in ancient customs, and it shows us the power of history, religion, ritual and tradition.

For Jews of course, the story is different. The Jewish explosion from Sicily was at its height in 1493 when the Spanish Inquisition reached the island.

But after 500 years, today there is a slight resurgence of the Jewish community when the Great Synagogue of Palermo was reopened in 2017. The building had been taken and used as a monastery, but it has now been returned to its historic owners.

The famous Godfather movies have two Sicilian weddings in the films. It is interesting and relevant to this discussion that they are very different. One is more modern and one very old-fashioned. One movie scholar writes that As noted in the screenplay, Michaels wedding is the same in feeling and texture as it might have been five hundred years ago, with all the ritual and pageantry, as it has always been, in Sicily.

Lengthy engagements are still the norm but that is mostly due to financial concerns. Having a good job to support a family and pay for a home leads to courtships as long and six or seven years. And when the couple finally announces their engagement, it could still be another year or two until the wedding. The brides family bears most of the costs of the wedding. There are no bridal showers are we know them, but they do now use the bridal registry, which helps the couple establish their home.

Divorce was uncommon until the 1970s. The 1961 film, Divorce, Italian Style, is based on the fact that divorce was actually illegal then, and the story involves a husband fantasizing about getting rid of his wife. Apparently a satire, or dark comedy, it plays with the concept that its OK to murder your wife. At least that is what I gather, I havent seen it.

Until about 1900 most Italian weddings, complete with dowries, were arranged by consent of the spouses' parents. A girl might be informally betrothed while fairly young, perhaps at 14, and wed at around the age of 18, although there were instances of girls marrying at 15. Sicilian marriages are no longer arranged by parents, and today your will even see public displays of affection and all the trappings of modern love and modern life. And today you can choose to have a civil ceremony, officiated by a mayor or civil registrar. Shocking! Same-sex civil unions and unregistered cohabitation have been legally recognized since June 2016.

Due to immigration and the exposure to the rest of the world, especially the EU and the U.S., the modern, western style wedding is in full swing. Even without travel opportunities, through TV and magazines, most Sicilians have adopted modern attitudes.

Lois Heckman is a certified Celebrant officiating in the Poconos and beyond. She writes about creating meaningful weddings, focusing on ceremony, ritual, and diverse traditions. Find her on Instagram, Facebook and website: http://www.LoisHeckman.com.

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The evolution of Sicilian weddings - Pocono Record

For Your Post-Holiday Enjoyment, Healthy ID Snacks – Discovery Institute

Yesterday, Evolution News offered appetizers for 2020. But maybe you are bothered already by overeating during the recent holidays. No problem. Here are healthy nuggets with an ID flavor to snack on.

Those plastic suction cups we use on glass fall off after a while. One would never think of getting them to stick to uneven rock. How about fastening them to rock underwater in a strong current? Impossible? The aquatic larvae of the net-winged midge do it. To the amazement of biologists from the University of Cambridge, these larvae even move around the rocks without losing their grip while subjected to absolutely enormous forces trying to pull them off, reports Phys.org.

The larvae have the ability to quickly detach and reattach to underwater rocks in torrential alpine rivers that can flow as fast as three metres per second. Their highly specialised suction organs are so strong that only forces over 600 times their body weight can detach them. [Emphasis added.]

The investigators had difficulty balancing while standing knee deep in the river, but they found the larvae grazing on the underwater rocks, apparently oblivious to the torrents bearing down on them. The larvae possess the highest attachment strength ever recorded in insects. Whats their secret?

The researchers found that a central piston, controlled by specific muscles, is used to create the suction and enable each larva to form a very tight seal with the surface of the rock. A dense array of tiny hairs come into contact with the rock surface, helping to keep the larva in place. When it needs to move, other muscles control a tiny slit on the suction disc, pulling the disc open to allow the suction organ to detach. This is the first time such an active detachment mechanism has been seen in any biological system.

Engineers could imitate this mechanism for numerous applications in industry and medicine.

The design inference is at work in the Astrophysical Journal. Astronomers are wondering about lights that appear and disappear in deep space. About a hundred anomalous sources have been detected. The New York Post says that scientists arent ruling out extraterrestrial intelligences as the cause. But of course, IDs design filter requires eliminating chance or nature first.

They say the blinking lights are most likely derived from natural, if somewhat extreme astrophysical sources, adding that the finding could change the study of astrophysics forever.

The implications of finding such objects extend from traditional astrophysics fields to the more exotic searches for evidence of technologically advanced civilizations, the authors write in their report, recently published in the Astronomical Journal.

Theyre not supernovae or any other known phenomena. So while intelligent design is barred from most journals, it remains useful. In fact, the researchers are very excited about looking for intelligent causes.

Archaea constitute a separate domain of life from bacteria, yet they, too, have outboard motors that help them swim. Phys.org describes their place in the living world:

Archaea, bacteria and eukaryotes are what biologists call the three domains of life. Of these three, archaea form an important link within the evolutionary theory. They are the direct ancestors of eukaryotes, but resemble bacteria in structure and organization. Archaea can colonize hot sulphur springs or extremely saline lakes, but can also be found in the ocean or in the human intestine and on the skin. Unlike bacteria, archaea have been relatively little researched because no pathogenic forms have been identified so far.

The outboard motor of some archaea is so different from the bacterial flagellum, it is called an archaellum instead. Like the flagellum, the archaellum appears to be irreducible complex. A paper in Nature Microbiology describes what happens when one part of this unique nanomachine is removed:

In Archaea, motility is mediated by the archaellum, a rotating type IV pilus-like structure that is a unique nanomachine for swimming motility in nature.

Strikingly, Sulfolobus cells that lack the S-layer component bound by FlaF assemble archaella but cannot swim. These collective results support a model where a FlaG filament capped by a FlaGFlaF complex anchors the archaellum to the S-layer to allow motility.

The authors note that Motility structures are vital in all three domains of life, and yet each type of machine is unique from those in the other domains. Phys.org adds:

It is important for microorganisms to be able to move actively so that if their environment deteriorates they can seek better living conditions. Bacteria use what is known as the flagellum, a complex structure requiring up to 50 proteins that assemble according to a strict timetable. Scientists assumed that archaea used the same structure as bacteria to swim from one place to another. But after sequencing the first archaeal genomes, the researchers discovered that archaea did not possess flagella operons. Instead, archaea swim using a structure called an archaellum. It consists of only seven subunits in the model organism Sulfolobus acidocaldarius used by Albers, which lives in highly acidic hydrothermal springs. Nevertheless, this relatively simple structure can perform the same functions as the bacterial flagellum, she explains.

The archaellum cannot be an ancestor of the flagellum, though, because its in a different domain of life, and uses different parts. If five parts in a mousetrap confirm intelligent design, seven subunits in the archaellum make an even stronger case. Undoubtedly the subunits have even more constituent parts.

So here, the most primitive microbes come already equipped with rotary outboard motors, and these tiny cells also live in some of the most extreme environments on the planet including hot springs at the boiling point. See our Archaea Have Their Own Rotary Propellers for more about the amazing archaellum that is profoundly different from the bacterial flagellum yet performs a similar function. Incidentally, it is also highly conserved in all archaeal species.

Youve had some ID cookies. Now here are some small candy pieces. Dont worry about the calories. Theyre all in your head!

The ultimate non-stick coating developed at McMaster University was inspired by the lotus leaf. Made into a plastic wrap, it stops dangerous superbugs in hospitals and food packaging.

Penn State announced a New, slippery toilet coating provides cleaner flushing, saves water. This biomimetic research has already led to products you can buy from SpotLESS Materials. The spray coating was inspired by the pitcher plant. Watch the video to see how well it works on glass and ceramics.

Soil has been called that thin layer on the planet that stands between us and starvation. Phys.org reported on work at the University of Wisconsin-Madison that shows how bacteria can degrade solid bedrock, jump-starting a long process of alteration that creates the mineral portion of soil. Bacteria speed up redox reactions that let them eat the rock because of a biological invention of a protein that allows cells to make electrical contact with minerals. Moreover, the bacteria couple the oxidation of iron to make ATP, the energy molecule in all forms of life. The way bacteria metabolize rock to begin soil formation has been going on basically forever, but unknown to us.

Better desalination machines and water filtration methods could be coming, thanks to mechanisms inspired by our own body cells. Aquaporins are essential membrane proteins that serve as water channels in the cells of our eyes, kidneys and other watery organs. Scientists at the University of Texas at Austin couldnt mimic aquaporins exactly, but found a way to create water wires that work a thousand times better than existing desalination systems.

It is difficult to even effectively mimic the complexities of how the human body works, especially at the molecular level, he said. This time, however, nature was the starting point for an even greater discovery than we could have ever hoped for.

Happy ID snacking all meat and no fat!

Image: Special repellent properties of the lotus leaf inspired the ultimate non-stick coating; by William Thielicke (more images here), [CC BY-SA 4.0], via Wikimedia Commons.

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For Your Post-Holiday Enjoyment, Healthy ID Snacks - Discovery Institute